Jansen M Andrew, Franz Nico M
School of Life Sciences, PO Box 874501, Arizona State University, Tempe, AZ 85287-4501.
Zookeys. 2015 Oct 20(528):1-133. doi: 10.3897/zookeys.528.6001. eCollection 2015.
This contribution adopts the taxonomic concept annotation and alignment approach. Accordingly, and where indicated, previous and newly inferred meanings of taxonomic names are individuated according to one specific source. Articulations among these concepts and pairwise, logically consistent alignments of original and revisionary classifications are also provided, in addition to conventional nomenclatural provenance information. A phylogenetic revision of the broad-nosed weevil genera Minyomerus Horn, 1876 sec. O'Brien & Wibmer (1982), and Piscatopus Sleeper, 1960 sec. O'Brien & Wibmer (1982) (Curculionidae [non-focal]: Entiminae [non-focal]: Tanymecini [non-focal]) is presented. Prior to this study, Minyomerus sec. O'Brien & Wibmer (1982) contained seven species, whereas the monotypic Piscatopus sec. O'Brien & Wibmer (1982) was comprised solely of Piscatopus griseus Sleeper, 1960 sec. O'Brien & Wibmer (1982). We thoroughly redescribe these recognized species-level entities and furthermore describe ten species as new to science: Minyomerus bulbifrons sec. Jansen & Franz (2015) (henceforth: [JF2015]), sp. n., Minyomerus aeriballux [JF2015], sp. n., Minyomerus cracens [JF2015], sp. n., Minyomerus gravivultus [JF2015], sp. n., Minyomerus imberbus [JF2015], sp. n., Minyomerus reburrus [JF2015], sp. n., Minyomerus politus [JF2015], sp. n., Minyomerus puticulatus [JF2015], sp. n., Minyomerus rutellirostris [JF2015], sp. n., and Minyomerus trisetosus [JF2015], sp. n. A cladistic analysis using 46 morphological characters of 22 terminal taxa (5/17 outgroup/ingroup) yielded a single most-parsimonious cladogram (L = 82, CI = 65, RI = 82). The analysis strongly supports the monophyly of Minyomerus [JF2015] with eight unreversed synapomorphies, and places Piscatopus griseus sec. O'Brien & Wibmer (1982) within the genus as sister to Minyomerus rutellirostris [JF2015]. Accordingly, Piscatopus sec. Sleeper (1960), syn. n. is changed to junior synonymy of Minyomerus [JF2015], and its sole member Piscatopus griseus sec. Sleeper (1960) is moved to Minyomerus [JF2015] as Minyomerus griseus [JF2015], comb. n. In addition, the formerly designated type Minyomerus innocuus Horn, 1876 sec. Pierce (1913), syn. n. is changed to junior synonymy of Minyomerus microps (Say, 1831) [JF2015] which has priority. The genus is widespread throughout western North America, ranging from Canada to Mexico and Baja California. Apparent patterns of interspecific diversity of exterior and genitalic morphology, varying host plant ranges, overlapping and widely extending species distributions, suggest an early origin for Minyomerus [JF2015], with a diversification that likely followed the development of North American desert biomes. Three species in the genus - i.e., Minyomerus languidus Horn, 1876 [JF2015], Minyomerus microps [JF2015], and Minyomerus trisetosus [JF2015] - are putatively considered parthenogenetic.
本论文采用分类学概念注释和比对方法。因此,在有指示的地方,分类学名称先前和新推断的含义根据一个特定来源进行区分。除了传统的命名来源信息外,还提供了这些概念之间的关联以及原始分类和修订分类的成对逻辑一致比对。本文对阔鼻象甲属(Minyomerus Horn,1876,隶属于O'Brien & Wibmer,1982)和鳞象甲属(Piscatopus Sleeper,1960,隶属于O'Brien & Wibmer,1982)(象甲科[非重点类群]:隐喙象亚科[非重点类群]:长角象族[非重点类群])进行了系统发育修订。在本研究之前,O'Brien & Wibmer(1982)的Minyomerus包含7个物种,而单型的Piscatopus(O'Brien & Wibmer,1982)仅由Piscatopus griseus Sleeper,1960(O'Brien & Wibmer,1982)组成。我们对这些已确认的物种级实体进行了全面重新描述,并另外描述了10个新物种:球额阔鼻象甲(Minyomerus bulbifrons [JF2015]),新种;气球阔鼻象甲(Minyomerus aeriballux [JF2015]),新种;惊鸣阔鼻象甲(Minyomerus cracens [JF2015]),新种;重负阔鼻象甲(Minyomerus gravivultus [JF2015]),新种;绒喙阔鼻象甲(Minyomerus imberbus [JF2015]),新种;糙刺阔鼻象甲(Minyomerus reburrus [JF2015]),新种;文雅阔鼻象甲(Minyomerus politus [JF2015]),新种;污斑阔鼻象甲(Minyomerus puticulatus [JF2015]),新种;槽喙阔鼻象甲(Minyomerus rutellirostris [JF2015]),新种;三鬃阔鼻象甲(Minyomerus trisetosus [JF2015]),新种。使用22个终端分类单元(5个外类群/17个内类群)的46个形态特征进行的分支分析产生了一个单一的最简约分支图(L = 82,CI = 65,RI = 82)。该分析有力地支持了具有8个不可逆共衍征的Minyomerus [JF2015]的单系性,并将Piscatopus griseus(O'Brien & Wibmer,1982)置于该属内,作为槽喙阔鼻象甲(Minyomerus rutellirostris [JF2015])的姐妹种。因此,Piscatopus(Sleeper,1960),新异名被改为Minyomerus [JF2015]的次异名,其唯一成员Piscatopus griseus(Sleeper,1960)作为灰阔鼻象甲(Minyomerus griseus [JF2015]),新组合被转移到Minyomerus [JF2015]。此外,先前指定的模式种无害阔鼻象甲(Minyomerus innocuus Horn,1876,隶属于Pierce,1913),新异名被改为具有优先权的微阔鼻象甲(Minyomerus microps (Say, 1831) [JF2015]的次异名。该属广泛分布于北美西部,从加拿大到墨西哥和下加利福尼亚。外部和生殖器形态的种间多样性、不同的寄主植物范围、重叠且广泛分布的物种分布的明显模式,表明Minyomerus [JF2015]起源较早,其多样化可能跟随北美沙漠生物群落的发展。该属中的三个物种——即慵懒阔鼻象甲(Minyomerus languidus Horn,1876 [JF2015])、微阔鼻象甲(Minyomerus microps [JF2015])和三鬃阔鼻象甲(Minyomerus trisetosus [JF2015])——被推定为孤雌生殖。