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新热带区竹节虫研究XX:来自法属圭亚那的一个新属和16个新物种。

Studies on neotropical Phasmatodea XX: A new genus and 16 new species from French Guiana.

作者信息

Conle Oskar V, Hennemann Frank H, Bellanger Yannick, Lelong Philippe, Jourdan Toni, Valero Pablo

机构信息

Am Freischütz 16, 47058 Duisburg, Germany..

出版信息

Zootaxa. 2020 Jul 14;4814(1):zootaxa.4814.1.1. doi: 10.11646/zootaxa.4814.1.1.

Abstract

The present paper describes 16 new species and one new genus from French Guiana and numerous taxonomic changes are proposed prior to the publication of a comprehensive guide to the Phasmatodea of French Guiana. The following 16 new species are described and illustrated: Phanocles procerus n. sp., Phanocloidea lobulatipes n. sp., Cladomorphus guianensis n. sp., Hirtuleius gracilis n. sp., Parastratocles rosanti n. sp., Parastratocles fuscomarginatus n. sp., Paraprisopus apterus n. sp., Paraprisopus multicolorus n. sp., Agrostia longicerca n. sp., Isagoras similis n. sp., Paragrostia brulei n. sp., Prexaspes globosicaput n. sp., Prexaspes guianensis n. sp., Dinelytron cahureli n. sp., Prisopus clarus n. sp. and Prisopus conocephalus n. sp.. The new genus Paragrostia n. gen. is established for the newly described Paragrostia brulei n. sp. and Paragrostia flavimaculata (Heleodoro, Mendes Rafael, 2017) n. comb. the latter of which is here transferred from Agrostia Redtenbacher, 1906.                Fifty-six new combinations are proposed with species transferred to other genera: Bacteria pallidenotata Redtenbacher, 1908, is transferred to Phanocloidea Zompro, 2001 (n. comb.); Bacteria maroniensis Chopard, 1911 is transferred to Phanocles Stål, 1875 (n. comb.); Cladomorphus gibbosus (Chopard, 1911) is transferred to Hirtuleius Stål, 1875 (n. comb.); Stratocles soror Redtenbacher, 1906, Parastratocles lugubris (Redtenbacher, 1906) and Parastratocles cryptochloris (Rehn, 1904) are transferred to Brizoides Redtenbacher, 1906 (n. comb.); Stratocles xanthomela (Olivier, 1792), Stratocles forcipatus Bolívar, 1896 and Stratocles tessulatus (Olivier, 1792) are transferred to Parastratocles (n. comb.); Olcyphides cinereus (Olivier, 1792), Perliodes affinis Redtenbacher, 1906, Perliodes nigrogranulosus Redtenbacher, 1906, Perliodes sexmaculatus Redtenbacher, 1906, Isagoras rugicollis (Gray, 1835), Isagoras sauropterus Rehn, 1947, Brizoides viridipes (Rehn, 1905) and Brizoides graminea Redtenbacher, 1906 are transferred to Agrostia Redtenbacher, 1906 (n. comb.); Agrostia flavimaculata Heleodoro, Mendes Rafael, 2017 is transferred to Paragrostia n. gen. (n. comb.); Isagoras affinis Chopard, 1911, Isagoras chocoensis Hebard, 1921, Isagoras metricus Rehn, 1947 and Isagoras schraderi Rehn, 1947 are transferred to Tenerella Redtenbacher, 1906 (n. comb.); Xerosoma glyptomerion Rehn, 1904 is transferred to Isagoras Stål, 1875 (n. comb.); Isagoras venosus (Burmeister, 1838), Paraphasma paulense Rehn, 1918 and Paraphasma quadratum (Bates, 1865) are transferred to Prexaspes Stål, 1875 (n. comb.); Prexaspes (Prexaspes) cneius (Westwood, 1859) is transferred to Tenerella Redtenbacher, 1906 (n. comb.); Prexaspes lateralis (Fabricius, 1775) is transferred to Paraphasma Redtenbacher, 1906 (n. comb.); Isagoras santara (Westwood, 1859) and Prexaspes olivaceus Chopard, 1911 are transferred to Periphloea Redtenbacher, 1906 (n. comb.); Dinelytron agrion Westwood, 1859 is transferred to Paraprisopus Redtenbacher, 1906 (n. comb.); Anarchodes atrophicus (Pallas, 1772) is transferred to Ignacia Rehn, 1904 (n. comb.); Planudes asperus Bellanger Conle, 2013, Planudes brunni Redtenbacher, 1906, Planudes cortex Hebard, 1919, Planudes crenulipes Rehn, 1904, Planudes funestus Redtenbacher, 1906, Planudes melzeri Piza, 1937, Planudes molorchus (Westwood, 1859), Planudes paxillus (Westwood, 1859), Planudes perillus Stål, 1875, Planudes pygmaeus (Redtenbacher, 1906) and Planudes taeniatus Piza, 1944 are transferred to Isagoras Stål, 1875 (n. comb.); Prisopoides atrobrunneus Heleodoro Rafael, 2020, Prisopoides brunnescens Heleodoro Rafael, 2020, Prisopoides caatingaensis Heleodoro Rafael, 2020 and Prisopoides villosipes (Redtenbacher, 1906) are transferred to Prisopus Peletier de Saint Fargeau Serville, 1828 (n. comb.); Melophasma antillarum (Caudell, 1914), Melophasma brachypterum Conle, Hennemann Gutiérrez, 2011, Melophasma colombianum Conle, Hennemann Gutiérrez, 2011 and Melophasma vermiculare Redtenbacher, 1906 are transferred to Paraprisopus Redtenbacher, 1906 (n. comb.); Prexaspes (Elasia) ambiguus (Stoll, 1813), Prexaspes (Elasia) brevipennis (Burmeister, 1838), Prexaspes (Elasia) pholcus (Westwood, 1859), Prexaspes (Elasia) viridipes Redtenbacher, 1906 and Prexaspes (Elasia) vittata (Piza, 1985) are transferred to Prexaspes Stål, 1875 (n. comb.).                Twenty-six new synonymies are established: Perliodes Redtenbacher, 1906 and Chlorophasma Redtenbacher, 1906 are synonymised with Agrostia Redtenbacher, 1906 (n. syn.); Chlorophasma Redtenbacher, 1906 is synonymised with Agrostia Redtenbacher, 1906 (n. syn.); Elasia Redtenbacher, 1906 is synonymised with Prexaspes Stål, 1875 (n. syn.); Prisopoides Heleodoro Rafael, 2020 is synonymised with Prisopus Peletier de Saint Fargeau Serville, 1828 (n. syn.); Melophasma Redtenbacher, 1906 is synonymised with Paraprisopus Redtenbacher, 1906 (n. syn.); Bacteria crassipes Chopard, 1911 is synonymised with Bacteria pallidenotata Redtenbacher, 1908 (n. syn.); Perliodes grisescens Redtenbacher, 1906 and Metriophasma (Metriophasma) pallidum (Chopard, 1911) are synonymised with Agrostia cinerea (Olivier, 1792) (n. syn.); Perliodes nigrogranulosus Redtenbacher, 1906 and Metriophasma (Metriophasma) ocellatum (Piza, 1937) are synonymised with Isagoras rugicollis (Gray, 1835) (n. syn.); Isagoras chopardi Hebard, 1933 is synonymised with Tenerella cneius (Westwood, 1859) (n. syn.); Isagoras proximus Redtenbacher, 1906 is synonymised with Isagoras glyptomerion (Rehn, 1904) (n. syn.); Chlorophasma hyalina Redtenbacher, 1906 is synonymised with Agrostia graminea (Redtenbacher, 1906) (n. syn.); Isagoras nitidus Redtenbacher, 1906 is synonymised with Anisa flavomaculatus (Gray, 1835) (n. syn.); Prexaspes acuticornis (Gray, 1835) is synonymised with Prexaspes servillei (Gray, 1835) (n. syn.); Prexaspes nigromaculatus Chopard, 1911 is synonymised with Periphloea santara (Westwood, 1859) (n. syn.); Prexaspes (Elasia) janus Kirby, 1904 is synonymised with Paraphasma maculatum (Gray, 1835) (n. syn.); Prexaspes dictys (Westwood, 1859) is synonymised with Prexaspes brevipennis (Burmeister, 1838) (n. syn.); Parastratocles aeruginosus Redtenbacher, 1906: 107 is synonymised with Parastratocles forcipatus Bolívar, 1896 (n. syn.); Parastratocles carbonarius (Redtenbacher, 1906: 106) is synonymised with Parastratocles lugubris (Redtenbacher, 1906) (n. syn.); Prisopus spinicollis Burmeister, 1838, Prisopus spiniceps Burmeister, 1838 and Prisopus cornutus Gray, 1835 are synonymised with Prisopus ohrtmanni (Lichtenstein, 1802) (n. syn.); the genus Planudes Stål, 1875 is synonymised with Isagoras Stål, 1875 (n. syn.); Pseudophasma annulipes (Redtenbacher, 1906) is synonymised with Pseudophasma blanchardi (Westwood, 1859) (n. syn.); Ignacia appendiculatum (Kirby, 1904) is synonymised with Anarchodes atrophicus (Pallas, 1772) (n. syn.).                Isagoras obscurum Guérin-Méneville, 1838 is shown to have been erroneously synonymised with Isagoras rugicollis (Gray, 1835) and is here re-established as a valid species (rev. stat.). Pseudophasma castaneum (Bates, 1865) is re-established as a valid species here (rev. stat.).                Paraprisopus Redtenbacher, 1906 and the entire tribe Paraprisopodini are transferred to Pseudophasmatidae: Pseudophasmatinae (n. comb.).                Lectotypes are designated for Perliodes grisescens Redtenbacher, 1906, Isagoras plagiatus Redtenbacher, 1906.Neotypes are designated for Agrostia cinerea (Olivier, 1792), Prexaspes ambiguus (Stoll, 1813), Prisopus horridus (Gray, 1835) and Prisopus sacratus (Olivier, 1792).

摘要

本文描述了来自法属圭亚那的16个新物种和1个新属,并在法属圭亚那竹节虫综合指南出版之前提出了许多分类学上的变动。描述并图示了以下16个新物种:长颈幻角竹节虫(Phanocles procerus n. sp.)、小叶幻角竹节虫(Phanocloidea lobulatipes n. sp.)、圭亚那枝形竹节虫(Cladomorphus guianensis n. sp.)、纤细赫氏竹节虫(Hirtuleius gracilis n. sp.)、罗桑蒂副层竹节虫(Parastratocles rosanti n. sp.)、褐缘副层竹节虫(Parastratocles fuscomarginatus n. sp.)、无翅副原竹节虫(Paraprisopus apterus n. sp.)、多色副原竹节虫(Paraprisopus multicolorus n. sp.)、长角禾叶竹节虫(Agrostia longicerca n. sp.)、似伊氏竹节虫(Isagoras similis n. sp.)、布氏副禾叶竹节虫(Paragrostia brulei n. sp.)、球头普氏竹节虫(Prexaspes globosicaput n. sp.)、圭亚那普氏竹节虫(Prexaspes guianensis n. sp.)、卡胡雷氏迪氏竹节虫(Dinelytron cahureli n. sp.)、透明普氏竹节虫(Prisopus clarus n. sp.)和锥头普氏竹节虫(Prisopus conocephalus n. sp.)。为新描述的布氏副禾叶竹节虫(Paragrostia brulei n. sp.)和黄斑副禾叶竹节虫(Paragrostia flavimaculata (Heleodoro, Mendes Rafael, 2017) n. comb.)建立了新属副禾叶竹节虫属(Paragrostia n. gen.),后者在此从1906年的禾叶竹节虫属(Agrostia Redtenbacher)转移而来。

提出了56个新组合,将物种转移到其他属:1908年的淡斑细菌竹节虫(Bacteria pallidenotata Redtenbacher)转移到2001年的幻角竹节虫属(Phanocloidea Zompro)(新组合);1911年的马罗尼细菌竹节虫(Bacteria maroniensis Chopard)转移到1875年的幻角竹节虫属(Phanocles Stål)(新组合);瘤状枝形竹节虫(Cladomorphus gibbosus (Chopard, 1911))转移到1875年的赫氏竹节虫属(Hirtuleius Stål)(新组合);2006年的姐妹层竹节虫(Stratocles soror Redtenbacher)、2006年的阴森副层竹节虫(Parastratocles lugubris (Redtenbacher, 1906))和2004年的隐绿副层竹节虫(Parastratocles cryptochloris (Rehn, 1904))转移到2006年的布里佐竹节虫属(Brizoides Redtenbacher)(新组合);1792年的黄斑层竹节虫(Stratocles xanthomela (Olivier, 1792))、1896年的钳状层竹节虫(Stratocles forcipatus Bolívar, 1896)和1792年的棋盘层竹节虫(Stratocles tessulatus (Olivier, 1792))转移到副层竹节虫属(Parastratocles)(新组合);1792年的灰绿奥氏竹节虫(Olcyphides cinereus (Olivier, 1792))、2006年的近缘佩氏竹节虫(Perliodes affinis Redtenbacher, 1906)、2006年的黑粒佩氏竹节虫(Perliodes nigrogranulosus Redtenbacher, 1906)、2006年的六斑佩氏竹节虫(Perliodes sexmaculatus Redtenbacher, 1906)、1835年的糙颈伊氏竹节虫(Isagoras rugicollis (Gray, 1835))、1947年的蜥翅伊氏竹节虫(Isagoras sauropterus Rehn, 1947)、2005年的绿足布里佐竹节虫(Brizoides viridipes (Rehn, 1905))和2006年的禾本科布里佐竹节虫(Brizoides graminea Redtenbacher, 1906)转移到1906年的禾叶竹节虫属(Agrostia Redtenbacher)(新组合);2017年的黄斑禾叶竹节虫(Agrostia flavimaculata Heleodoro, Mendes Rafael, 2017)转移到副禾叶竹节虫属(Paragrostia n. gen.)(新组合);2011年的近缘伊氏竹节虫(Isagoras affinis Chopard, 1911)、1921年的乔科伊氏竹节虫(Isagoras chocoensis Hebard, 1921)、1947年的米氏伊氏竹节虫(Isagoras metricus Rehn, 1947)和1947年的施氏伊氏竹节虫(Isagoras schraderi Rehn, 1947)转移到2006年的特内雷拉竹节虫属(Tenerella Redtenbacher)(新组合);2004年的刻纹干竹节虫(Xerosoma glyptomerion Rehn, 1904)转移到1875年的伊氏竹节虫属(Isagoras Stål)(新组合);1838年的脉纹伊氏竹节虫(Isagoras venosus (Burmeister, 1838))、1918年的保罗副竹节虫(Paraphasma paulense Rehn, 1918)和1865年的方形副竹节虫(Paraphasma quadratum (Bates, 1865))转移到1875年的普氏竹节虫属(Prexaspes Stål)(新组合);1859年的克氏普氏竹节虫(Prexaspes (Prexaspes) cneius (Westwood, 1859))转移到2006年的特内雷拉竹节虫属(Tenerella Redtenbacher)(新组合);1775年的侧斑普氏竹节虫(Prexaspes lateralis (Fabricius, 1775))转移到1906年的副竹节虫属(Paraphasma Redtenbacher)(新组合);1859年的桑塔拉伊氏竹节虫(Isagoras santara (Westwood, 1859))和1911年的橄榄色普氏竹节虫(Prexaspes olivaceus Chopard, 1911)转移到2006年的周韧竹节虫属(Periphloea Redtenbacher)(新组合);1859年的食蚜迪氏竹节虫(Dinelytron agrion Westwood, 1859)转移到1906年的副原竹节虫属(Paraprisopus Redtenbacher)(新组合);1772年的萎缩无秩序竹节虫(Anarchodes atrophicus (Pallas, 1772))转移到2004年的伊格纳西亚竹节虫属(Ignacia Rehn)(新组合);2013年的粗糙普拉努德竹节虫(Planudes asperus Bellanger Conle)、1906年的棕色普拉努德竹节虫(Planudes brunni Redtenbacher)、1919年的树皮普拉努德竹节虫(Planudes cortex Hebard)、1904年的具细圆齿足普拉努德竹节虫(Planudes crenulipes Rehn)、1906年的不祥普拉努德竹节虫(Planudes funestus Redtenbacher)、1937年的梅尔泽普拉努德竹节虫(Planudes melzeri Piza)、1859年的莫洛赫普拉努德竹节虫(Planudes molorchus (Westwood, 1859))、1859年的小垫普拉努德竹节虫(Planudes paxillus (Westwood, 1859))、1875年的危险普拉努德竹节虫(Planudes perillus Stål)、1906年的侏儒普拉努德竹节虫(Planudes pygmaeus (Redtenbacher, 1906))和1944年的带纹普拉努德竹节虫(Planudes taeniatus Piza)转移到1875年的伊氏竹节虫属(Isagoras Stål)(新组合);2020年的黑褐拟普氏竹节虫(Prisopoides atrobrunneus Heleodoro Rafael)、2020年的浅褐拟普氏竹节虫(Prisopoides brunnescens Heleodoro Rafael)、2020年的卡廷加拟普氏竹节虫(Prisopoides caatingaensis Heleodoro Rafael)和1906年的具毛足拟普氏竹节虫(Prisopoides villosipes (Redtenbacher, 1906))转移到1828年的普氏竹节虫属(Prisopus Peletier de Saint Fargeau Serville)(新组合);2014年的安的列斯岛变色竹节虫(Melophasma antillarum (Caudell, 1914))、2011年的短翅变色竹节虫(Melophasma brachypterum Conle, Hennemann Gutiérrez)、2011年的哥伦比亚变色竹节虫(Melophasma colombianum Conle, Hennemann Gutiérrez)和1906年的蠕虫状变色竹节虫(Melophasma vermiculare Redtenbacher)转移到1906年的副原竹节虫属(Paraprisopus Redtenbacher)(新组合);1813年的模糊普氏竹节虫(Prexaspes (Elasia) ambiguus (Stoll, 1813))、1838年的短翅普氏竹节虫(Prexaspes (Elasia) brevipennis (Burmeister, 1838))、1859年的幽灵普氏竹节虫(Prexaspes (Elasia) pholcus (Westwood, 185

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