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东南亚和新几内亚螽斯科(直翅目,硕螽亚目)的新分类群及记录,并附属检索表

New taxa and records of Gryllacrididae (Orthoptera, Stenopelmatoidea) from South East Asia and New Guinea with a key to the genera.

作者信息

Ingrisch Sigfrid

机构信息

Zoologisches Forschungsmuseum Alexander Koenig, Adenauerallee 160, D- 53113 Bonn, Germany..

出版信息

Zootaxa. 2018 Nov 5;4510(1):1-278. doi: 10.11646/zootaxa.4510.1.1.

Abstract

New records of Gryllacrididae from South East Asia to New Guinea are reported, new taxa are described and type specimens from European collections redescribed. The wing venation of the tegmina and its variation within the family is discussed. A new key to the 50 genera and subgenera of Gryllacrididae occurring in the area from South East Asia to New Guinea is provided. Six genera and two subgenera are introduced as new: Angustogryllacris gen. nov., Minigryllacris gen. nov., Monseremus gen. nov., Siamgryllacris gen. nov., Paraneanias gen. nov., Plexigryllacris gen. nov., Pseudasarca subgen. nov. of Asarcogryllacris Karny, 1937, Xiphilarnaca subgen. nov. of Metriogryllacris Karny, 1937. Two genera Gigantogryllacris Karny, 1937 and Pardogryllacris Karny, 1937 are reduced to subgeneric status under Gryllacris Audinet-Serville, 1831 and one monotypic genus Cyanogryllacris Karny, 1937 is synonymised with Capnogryllacris Karny, 1937.        70 species and nine subspecies are described as new: Eugryllacris gandaki sp. nov., Eugryllacris guomashan sp. nov., Eugryllacris crassicauda sp. nov., Eugryllacris crassicauda cambodiana ssp. nov., Eugryllacris serricauda sp. nov., Eugryllacris trabicauda sp. nov., Eugryllacris vermicauda sp. nov., Eugryllacris sulcata sp. nov., Eugryllacris inversa sp. nov., Gryllacris incornuta sp. nov., Gryllacris sok sp. nov., Gryllacris (Pardogryllacris) ovulicauda sp. nov., Gryllacris (Gigantogryllacris) bilineata sp. nov., Gryllacris (Gigantogryllacris) ligulata sp. nov., Lyperogryllacris forcipata sp. nov., Lyperogryllacris ocellata sp. nov., Lyperogryllacris khuntan sp. nov., Ocellarnaca disjuncta sp. nov., Ocellarnaca fusca sp. nov., Otidiogryllacris lawang sp. nov., Otidiogryllacris bamusbama sp. nov., Prosopogryllacris gamta sp. nov., Prosopogryllacris nigra sp. nov., Prosopogryllacris silacea sp. nov., Xanthogryllacris lineata sp. nov., Xanthogryllacris punctata sp. nov., Xanthogryllacris ralum sp. nov., Xanthogryllacris subrecta sp. nov., Aancistroger inarmatus sp. nov., Angustogryllacris bibulbata sp. nov., Aphanogryllacris sinustylata sp. nov., Aphanogryllacris nigritibiae sp. nov., Asarcogryllacris (Asarcogryllacris) brevis sp. nov., Asarcogryllacris (Asarcogryllacris) parapat sp. nov., Asarcogryllacris (Asarcogryllacris) robusta sp. nov., Asarcogryllacris (Pseudolarnaca) cornualis sp. nov., Asarcogryllacris (Pseudasarca) arborea sp. nov., Australogryllacris guttata sp. nov., Capnogryllacris (C.) erythrocephala fuscifrons ssp. nov., Capnogryllacris (C.) varifrons sp. nov., Capnogryllacris (C.) nigromaculata sp. nov., Capnogryllacris (C.) sakaerat toxica ssp. nov., Celebogryllacris brevitegmina sp. nov., Diaphanogryllacris annamita tenera ssp. nov., Diaphanogryllacris opulenta sp. nov., Diaphanogryllacris recta sp. nov., Diaphanogryllacris sinuata sp. nov., Eremus tigris sp. nov., Furcilarnaca chiangdao sp. nov., Furcilarnaca salit sp. nov., Furcilarnaca trilobata sp. nov., Haplogryllacris bilobulata sp. nov., Homogryllacris armigera sp. nov., Homogryllacris stabilis sp. nov., Larnaca (Larnaca) nigricornis sp. nov., Larnaca (Larnaca) tenuis sp. nov., Larnaca (Larnaca) samkos sp. nov., Larnaca (Larnaca) squamiptera sp. nov., Larnaca (Larnaca) subaptera sp. nov., Melaneremus sikkimensis sp. nov., Minigryllacris perpusilla sp. nov., Monseremus appendiculatus sp. nov., Neanias virens sp. nov., Neolarnaca vera nigrinotum ssp. nov., Phryganogryllacris extensa sp. nov., Phryganogryllacris gialaiensis fovealis ssp. nov., Phryganogryllacris nonangulata sp. nov., Siamgryllacris rufa sp. nov., Ultragryllacris pulchra nan ssp. nov., Ultragryllacris triangula sp. nov., Woznessenskia ampliata sp. nov., Woznessenskia bavi sp. nov., Zalarnaca (Glolarnaca) elegantula sp. nov., Zalarnaca (Glolarnaca) globiceps minor ssp. nov., Zalarnaca (Zalarnaca) maninjau sp. nov., Papuogryllacris diluta baiteta ssp. nov., Papuogryllacris rugifrons sp. nov., Papuogryllacris ligata bundi ssp. nov., Paraneanias striatus sp. nov., Plexigryllacris megastyla sp. nov.        The following nomenclatural changes had to be done: Gryllacris bancana Karny, 1930a stat. nov. is raised to full species from subspecies of Gryllacris obscura Brunner von Wattenwyl, 1888; Xanthogryllacris inquinata (Karny, 1928c) comb. stat. nov. is raised to full species from subspecies under Pardogryllacris dyak (Griffini, 1909); Eremus oberthuri Griffini, 1913a stat. nov. is raised to full species from subspecies under Eremus rugosifrons Brunner von Wattenwyl, 1888; Prosopogryllacris nigrovenosa (Karny, 1928b) stat. nov. is raised to full species from subspecies under Prosopogryllacris horvathi (Griffini, 1909); Xanthogryllacris punctipennis aurantiaca (Brunner von Wattenwyl, 1888) stat. reest. and Xanthogryllacris punctipennis dempwolffi (Griffini, 1909b) stat. reest. are treated as valid subspecies under Xanthogryllacris punctipennis (Walker, 1869) instead of being synonyms of the latter; Gryllacris thailandi Gorochov, 2007 and Gryllacris thailandi facemarmiger Gorochov et al., 2015 are downgraded to subspecies of Gryllacris vittata Walker, 1869. Gryllacris kledangensis Karny, 1923 and Gryllacris lombokiana Karny, 1926a become new synonyms of Gryllacris peracca Karny, 1923; Brachyntheisogryllacris abbreviata evolutior (Griffini, 1909a) becomes a new synonym of Brachyntheisogryllacris abbreviata (Brunner von Wattenwyl, 1888), and Haplogryllacris hieroglyphicoides (Chopard, 1924) a new synonym of Haplogryllacris simplex (Walker, 1871).        The following new combinations are proposed: Capnogryllacris (C.) basaliatrata (Griffini, 1909c) comb. nov. from Cyanogryllacris Karny, 1937; Haplogryllacris aliena Walker, 1869 comb. nov. and Haplogryllacris durgensis (Gupta Chandra, 2017) comb. nov. from Gryllacris Audinet-Serville, 1831; Homogryllacris artinii (Griffini, 1913b) comb. nov., Homogryllacris buyssoniana (Griffini, 1912c) comb. nov., Homogryllacris kurseonga (Griffini, 1913b) stat. comb. nov. and Homogryllacris ? maindroni (Griffini, 1913a) comb. nov. from Brachyntheisogryllacris Karny, 1937; Larnaca (Larnaca) eugenii (Griffini, 1914b) comb. nov. from Metriogryllacris Karny, 1937; Larnaca (Larnaca) larnacoides (Karny, 1937) comb. nov. and Larnaca (Larnaca) jacobsoni (Griffini, 1913c) comb. nov. from Melaneremus Karny, 1937.

摘要

报道了东南亚至新几内亚地区蟋螽科的新记录,描述了新分类单元,并重新描述了欧洲馆藏的模式标本。讨论了蟋螽前翅脉序及其在科内的变异。提供了一个新的检索表,用于鉴定东南亚至新几内亚地区的50个蟋螽属和亚属。引入了6个新属和2个新亚属:窄蟋螽属Angustogryllacris gen. nov.、小蟋螽属Minigryllacris gen. nov.、蒙氏蟋螽属Monseremus gen. nov.、暹罗蟋螽属Siamgryllacris gen. nov.、副尼蟋螽属Paraneanias gen. nov.、多突蟋螽属Plexigryllacris gen. nov.;1937年卡尔尼的粗角蟋螽属Asarcogryllacris的伪粗角亚属Pseudasarca subgen. nov.;1937年卡尔尼的中蟋螽属Metriogryllacris的剑突亚属Xiphilarnaca subgen. nov.。将1937年卡尔尼的巨蟋螽属Gigantogryllacris和帕氏蟋螽属Pardogryllacris降为1831年奥迪内-塞尔维的蟋螽属Gryllacris的亚属;将1937年卡尔尼的单型属蓝蟋螽属Cyanogryllacris与1937年卡尔尼的烟蟋螽属Capnogryllacris同义。描述了70个新物种和9个新亚种:甘达基真蟋螽Eugryllacris gandaki sp. nov.、郭马山真蟋螽Eugryllacris guomashan sp. nov.、粗尾真蟋螽Eugryllacris crassicauda sp. nov.、柬埔寨粗尾真蟋螽Eugryllacris crassicauda cambodiana ssp. nov.、锯尾真蟋螽Eugryllacris serricauda sp. nov.、横尾真蟋螽Eugryllacris trabicauda sp. nov.、蠕虫尾真蟋螽Eugryllacris vermicauda sp. nov.、沟尾真蟋螽Eugryllacris sulcata sp. nov.、内凹真蟋螽Eugryllacris inversa sp. nov.、无角蟋螽Gryllacris incornuta sp. nov.、索克蟋螽Gryllacris sok sp. nov.、(帕氏蟋螽亚属)卵尾蟋螽Gryllacris (Pardogryllacris) ovulicauda sp. nov.、(巨蟋螽亚属)双线蟋螽Gryllacris (Gigantogryllacris) bilineata sp. nov.、(巨蟋螽亚属)舌状蟋螽Gryllacris (Gigantogryllacris) ligulata sp. nov.、钳状跃蟋螽Lyperogryllacris forcipata sp. nov.、眼状跃蟋螽Lyperogryllacris ocellata sp. nov.、昆坦跃蟋螽Lyperogryllacris khuntan sp. nov.、间断眼蟋螽Ocellarnaca disjuncta sp. nov.、深色眼蟋螽Ocellarnaca fusca sp. nov.、拉旺耳蟋螽Otidiogryllacris lawang sp. nov.、巴姆斯巴耳蟋螽Otidiogryllacris bamusbama sp. nov.、加姆塔脸蟋螽Prosopogryllacris gamta sp. nov.、黑色脸蟋螽Prosopogryllacris nigra sp. nov.、粉质脸蟋螽Prosopogryllacris silacea sp. nov.、线纹黄蟋螽Xanthogryllacris lineata sp. nov.、斑点黄蟋螽Xanthogryllacris punctata sp. nov.、拉卢姆黄蟋螽Xanthogryllacris ralum sp. nov.、近直黄蟋螽Xanthogryllacris subrecta sp. nov.、无武装钩蟋螽Aancistroger inarmatus sp. nov.、双球窄蟋螽Angustogryllacris bibulbata sp. nov.、波状突隐蟋螽Aphanogryllacris sinustylata sp. nov.、黑胫隐蟋螽Aphanogryllacris nigritibiae sp. nov.、(粗角蟋螽亚属)短粗角蟋螽Asarcogryllacris (Asarcogryllacris) brevis sp. nov.、(粗角蟋螽亚属)邻接粗角蟋螽Asarcogryllacris (Asarcogryllacris) parapat sp. nov.、(粗角蟋螽亚属)粗壮粗角蟋螽Asarcogryllacris (Asarcogryllacris) robusta sp. nov.、(伪拉氏蟋螽亚属)角状粗角蟋螽Asarcogryllacris (Pseudolarnaca) cornualis sp. nov.、(伪粗角亚属)树栖粗角蟋螽Asarcogryllacris (Pseudasarca) arborea sp. nov.、斑点澳蟋螽Australogryllacris guttata sp. nov.、(烟蟋螽属)红额烟蟋螽棕额亚种Capnogryllacris (C.) erythrocephala fuscifrons ssp. nov.、(烟蟋螽属)杂色烟蟋螽Capnogryllacris (C.) varifrons sp. nov.、(烟蟋螽属)黑斑烟蟋螽Capnogryllacris (C.) nigromaculata sp. nov.、(烟蟋螽属)萨凯拉特毒烟蟋螽亚种Capnogryllacris (C.) sakaerat toxica ssp. nov.、短翅塞莱蟋螽Celebogryllacris brevitegmina sp. nov.、(透蟋螽属)安南透蟋螽柔弱亚种Diaphanogryllacris annamita tenera ssp. nov.、(透蟋螽属)华丽透蟋螽Diaphanogryllacris opulenta sp. nov.、(透蟋螽属)直透蟋螽Diaphanogryllacris recta sp. nov.、(透蟋螽属)波状透蟋螽Diaphanogryllacris sinuata sp. nov.、虎纹沙蟋螽Eremus tigris sp. nov.、清道叉蟋螽Furcilarnaca chiangdao sp. nov.、萨利叉蟋螽Furcilarnaca salit sp. nov.、三叶叉蟋螽Furcilarnaca trilobata sp. nov.、双叶单蟋螽Haplogryllacris bilobulata sp. nov.、武装同蟋螽Homogryllacris armigera sp. nov.、稳定同蟋螽Homogryllacris stabilis sp. nov.、(拉氏蟋螽属)黑角拉氏蟋螽Larnaca (Larnaca) nigricornis sp. nov.、(拉氏蟋螽属)细拉氏蟋螽Larnaca (Larnaca) tenuis sp. nov.、(拉氏蟋螽属)桑科斯拉氏蟋螽Larnaca (Larnaca) samkos sp. nov.、(拉氏蟋螽属)鳞翅拉氏蟋螽Larnaca (Larnaca) squamiptera sp. nov.、(拉氏蟋螽属)近无翅拉氏蟋螽Larnaca (Larnaca) subaptera sp. nov.、锡金黑蟋螽Melaneremus sikkimensis sp. nov.、极小小蟋螽Minigryllacris perpusilla sp. nov.、具附器蒙氏蟋螽Monseremus appendiculatus sp. nov.、绿色新尼蟋螽Neanias virens sp. nov.、(新拉氏蟋螽属)真正新拉氏蟋螽黑背亚种Neolarnaca vera nigrinotum ssp. nov.、扩展裸蟋螽Phryganogryllacris extensa sp. nov.、(裸蟋螽属)嘉莱裸蟋螽凹窝亚种Phryganogryllacris gialaiensis fovealis ssp. nov.、无角裸蟋螽Phryganogryllacris nonangulata sp. nov.、红色暹罗蟋螽Siamgryllacris rufa sp. nov.、(超蟋螽属)美丽超蟋螽矮小亚种Ultragryllacris pulchra nan ssp. nov.、(超蟋螽属)三角超蟋螽Ultragryllacris triangula sp. nov.、扩大沃兹蟋螽Woznessenskia ampliata sp. nov.、巴维沃兹蟋螽Woznessenskia bavi sp. nov.、(扎氏蟋螽属)优雅扎氏蟋螽Zalarnaca (Glolarnaca) elegantula sp. nov.、(扎氏蟋螽属)球头扎氏蟋螽小头亚种Zalarnaca (Glolarnaca) globiceps minor ssp. nov.、(扎氏蟋螽属)马宁贾岛氏蟋螽Zalarnaca (Zalarnaca) maninjau sp. nov.、(巴布亚蟋螽属)淡色巴布亚蟋螽拜泰塔亚种Papuogryllacris diluta baiteta ssp. nov.、(巴布亚蟋螽属)皱额巴布亚蟋螽Papuogryllacris rugifrons sp. nov.、(巴布亚蟋螽属)带状巴布亚蟋螽邦迪亚种Papuogryllacris ligata bundi ssp. nov.、具条纹副尼蟋螽Paraneanias striatus sp. nov.、大花柱多突蟋螽Plexigryllacris megastyla sp. nov.。进行了以下命名变更:1930a年卡尔尼的邦卡蟋螽Gryllacris bancana从1888年布伦纳·冯·瓦滕威尔的暗蟋螽Gryllacris obscura的亚种提升为独立物种;1928c年卡尔尼的污黄蟋螽Xanthogryllacris inquinata从1909年格里菲尼的达雅克帕氏蟋螽Pardogryllacris dyak的亚种提升为独立物种;1913a年格里菲尼的奥氏沙蟋螽Eremus oberthuri从1888年布伦纳·冯·瓦滕威尔的皱额沙蟋螽Eremus rugosifrons的亚种提升为独立物种;1928b年卡尔尼的黑脉脸蟋螽Prosopogryllacris nigrovenosa从1909年格里菲尼的霍氏脸蟋螽Prosopogryllacris horvathi的亚种提升为独立物种;1888年布伦纳·冯·瓦滕威尔的黄斑黄蟋螽Xanthogryllacris punctipennis aurantiaca和1909b年格里菲尼的黄斑黄蟋螽Xanthogryllacris punctipennis dempwolffi被重新认定为186

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