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细尺度地形对伊朗北部成熟山毛榉(Fagus orientalis Lipsky.)林树种直径空间分布的影响。

Fine-scale topographic influence on the spatial distribution of tree species diameter in old-growth beech (Fagus orientalis Lipsky.) forests, northern Iran.

机构信息

UCD Forestry, School of Agriculture and Food Science, University College Dublin, Dublin, Ireland.

Faculty of Natural Resources & Marine Sciences, Tarbiat Modares University, 46417-76489, Noor, Mazandaran, Iran.

出版信息

Sci Rep. 2022 May 10;12(1):7633. doi: 10.1038/s41598-022-10606-0.

Abstract

The Hyrcanian forest in northern Iran is threatened by human use and encroachment and has suffered degradation in some areas. The forest has been declared a World Heritage Site and management in the region is shifting from timber production to conservation. There is considerable interest in developing a greater understanding of these diverse forest communities to inform forest management and multiple use plans to maintain the diversity and resilience of these forests. The Hyrcanian forest is characterized by a complex topography of catenas ranging up mountain slopes. Topographic gradients greatly influence microhabitat conditions which in turn impact tree distribution. To date there has been limited research on the impacts of this diverse topography on the spatial distribution of tree species and tree diameters in Hyrcanian forests. Such information is necessary to better understand the regional traits of tree diameters in these natural mixed temperate forests before forest management occurs. We examined the influence of the area's catena topography on the spatial pattern of tree species and on species stand structure in terms of tree diameter distribution. To quantify these dynamics, we conducted a complete enumeration inventory of all trees with dbh >12 cm within a 7.947 ha study area that included three C-shaped (concave) and three V-shaped (convex) catenas. Geostatistical variogram analysis and Clark and Evans aggregation index were utilized to study the spatial distribution of tree diameters. Beech, alder, hornbeam, linden and Persian maple exhibited clustered patterns, and sour cherry, ash, and oak exhibited random patterns. Geostatistical analysis clearly revealed the substantial influence of catena topography on the diameter distributions of alder and linden, more subtle influence on the diameter distributions of beech, and a possible influence on Persian maple, providing valuable insight into stand structure over neighborhood-based indices alone. Alder and linden both exhibited strong spatial structure in their diameter distributions (56% and 86%, respectively) where their diameter was strongly correlated with trees within 108 m and 83 m, respectively, sharing more similar diameters to each other than trees beyond that distance. Beech, maple, and hornbeam exhibited very weak if any spatial structure over short distances. These findings can be used to support the alignment of forest management practices in managed Hyrcanian forests with goals of protecting and maintaining biodiversity and sustainable forest ecosystems, and to inform geospatial modeling of species diameter distributions in areas where a complete stem-map is not feasible.

摘要

伊朗北部的赫卡尼亚森林受到人类利用和侵占的威胁,一些地区已经退化。该森林已被宣布为世界遗产地,该地区的管理正从木材生产转向保护。人们对深入了解这些不同的森林群落产生了浓厚的兴趣,以便为森林管理和多用途规划提供信息,以维持这些森林的多样性和恢复力。赫卡尼亚森林的特点是山脉斜坡上有一系列复杂的阶地。地形梯度极大地影响了微生境条件,进而影响了树木的分布。迄今为止,关于这种多样的地形对赫卡尼亚森林树种和树木直径的空间分布的影响的研究有限。在进行森林管理之前,这种信息对于更好地了解这些天然混交温带森林的区域特征很有必要。我们研究了该地区阶地地形对树种空间格局以及树木直径分布方面的树种立地结构的影响。为了量化这些动态,我们对一个 7.947 公顷的研究区域内所有 dbh 大于 12 厘米的树木进行了全面的清点,该区域包括三个 C 形(凹形)和三个 V 形(凸形)阶地。我们利用地质统计学变差函数分析和克拉克和埃文斯聚集指数来研究树木直径的空间分布。山毛榉、桤木、榉木、椴树和枫香表现出聚集模式,而酸樱桃、白蜡和栎树表现出随机模式。地质统计学分析清楚地表明,阶地地形对桤木和椴树的直径分布有很大的影响,对山毛榉的直径分布有更微妙的影响,对枫香的直径分布可能有影响,为仅基于基于邻域的指数的林分结构提供了有价值的见解。桤木和椴树的直径分布都表现出很强的空间结构(分别为 56%和 86%),它们的直径与各自 108 米和 83 米范围内的树木强烈相关,彼此之间的直径比与更远距离的树木更相似。山毛榉、枫香和榉木在短距离内表现出非常弱的空间结构,如果有的话。这些发现可用于支持赫卡尼亚管理森林中的森林管理实践与保护和维持生物多样性和可持续森林生态系统的目标保持一致,并为在没有完整茎图的情况下提供物种直径分布的地理空间建模信息。

https://cdn.ncbi.nlm.nih.gov/pmc/blobs/63ec/9090739/42c2debd547f/41598_2022_10606_Fig1_HTML.jpg

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