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Physiol Behav. 1986;36(3):445-9. doi: 10.1016/0031-9384(86)90313-6.
Planaria Dugesia dorotocephala were bilaterally enucleated, decapitated caudal to the auricles, or sectioned at the level of the pharynx. None of these worms either altered or elicited any movement responses to stimulation by horizontal or overhead light even though they made such responses to mechanical stimulation. By comparison, all intact planaria responded vigorously to light stimulation but not to an elevation of water temperature. These results suggest that light receptors in these planaria might have evolved away from the body surface and are located in the ocelli.
将三角涡虫双侧摘除眼球,在耳状部后方断头,或在咽水平处切断。这些涡虫中没有一只对水平或头顶光刺激有任何改变或引发的运动反应,尽管它们对机械刺激会做出这样的反应。相比之下,所有完整的涡虫对光刺激反应强烈,但对水温升高没有反应。这些结果表明,这些涡虫中的光感受器可能已经从体表进化出来,位于眼点中。