Honorary Research Associate; Agriculture and Agri-Food Canada; Canadian National Collection of Insects; Arachnids and Nematodes; K. W. Neatby Bldg.; 960 Carling Avenue; Ottawa; Ontario; CANADA; K1A 0C6.
Al. I. Cuza University; Faculty of Biology; Bd. Carol I; Nr 11; 700506 Iasi; ROMANIA.
Zootaxa. 2023 Jul 19;5318(2):195-216. doi: 10.11646/zootaxa.5318.2.2.
Leptoomidae Gibson fam. nov. (Chalcidoidea) is described for the Eocene Baltic amber fossil genera Leptoomus Gibson, type genus, reassigned from Tanaostigmatidae, and Neanaperiallus Gibson, reassigned from Neanastatinae (Chalcidoidea: Eupelmidae) sensu Gibson (2009). One new species of Neanaperiallus, N. defunctus Fusu sp. nov., is described. The new family is differentiated from other families of Chalcidoidea that are partly characterized by a greatly enlarged acropleuron. In species of Leptoomidae the prepectus is anteriorly rounded to angulate and extends to or slightly over the posterolateral margin of the pronotum, with the dorsal prepectal margin intersecting the base of the tegula distinctly anterior to and forming an almost right-angle with the posterior margin of prepectus, and the posterior margin truncate along the anterior margin of the acropleuron. This prepectal structure is similar to that in Tanaostigmatidae and Cynipencyrtidae, except the prepectus is elongated anteriorly exterior to the pronotum in Tanaostigmatidae and interior to the lateral surface of the pronotum in Cynipencyrtidae. A difference in prepectal structure also indicates that an anteriorly elongated mesoscutal process internal to the pronotum in Encyrtidae is convergent to that of Cynipencyrtidae, and similarity in shape of the prepectus among Encyrtidae, Eopelma Gibson and Neanastatus Girault might be functionally correlated with an anterior elongation of the mesoscutal process. New or corrected morphological data are provided for the two included genera. Of other Eocene fossil genera originally classified in Neanastatinae, Brevivulva Gibson and Propelma Trjapitzin, are assigned to Neanastatidae sensu Burks et al. (2022) based on similar mesoscutellar structures. Possible relationships of Aspidopleura Gibson, a taxon with a puzzling combination of features, are discussed. Because Aspidopleura cannot be placed with confidence in any extinct or extant higher taxon it is treated as incertae sedis at family level within Chalcidoidea.
新分类单元 Leptoomidae Gibson, fam. nov. (膜翅目:细蜂总科),描述了始新世波罗的海琥珀化石属 Leptoomus Gibson,模式属,从 Tanaostigmatidae 重新分类,以及 Neanaperiallus Gibson,从 Neanastatinae (膜翅目:小蜂科)重新分类( Gibson 2009)。描述了一个 Neanaperiallus 的新种,N. defunctus Fusu sp. nov.。新科与其他部分特征为前盾片极度扩大的细蜂总科科之间存在差异。在 Leptoomidae 物种中,前胸前侧圆到有角,延伸到或略超过前胸的后外侧缘,背前胸前侧缘与盾片基部的交点明显在前部与前胸前侧缘形成几乎直角,后缘沿前盾片前缘截断。这种前胸前结构与 Tanaostigmatidae 和 Cynipencyrtidae 相似,但在 Tanaostigmatidae 中,前胸前侧在前胸外侧伸长,而在 Cynipencyrtidae 中,前胸前侧在前胸外侧表面内部伸长。前胸前结构的差异也表明,在小蜂科中,前胸内部向前伸长的中胸盾片与 Cynipencyrtidae 趋同,而在小蜂科、 Eopelma Gibson 和 Neanastatus Girault 中,前胸前的形状相似可能与中胸盾片的向前伸长有关。为两个包含的属提供了新的或更正的形态学数据。其他始新世化石属,Brevivulva Gibson 和 Propelma Trjapitzin,最初被归类于 Neanastatinae,根据相似的中胸盾片结构,被归类于 Neanastatidae sensu Burks et al. (2022)。讨论了具有令人费解特征组合的分类单元 Aspidopleura Gibson 的可能关系。由于 Aspidopleura 不能自信地归入任何已灭绝或现存的高级分类群,因此在膜翅目内被视为未确定科的地位。