Affanni J M, García Samartino L, Casanave E B, Dezi R
Respir Physiol. 1987 Feb;67(2):239-45. doi: 10.1016/0034-5687(87)90044-2.
The armadillo Chaetophractus villosus shows a remarkable respiratory adaptation when the nostrils are completely covered by soil: it is able to maintain efficient respiratory movements with the corresponding filling of the lungs, thanks to a mechanism that allows it to use the air filling the space between the soil particles, without invasion of the nasal cavities by those particles.