Kröger R H, Campbell M C, Fernald R D, Wagner H J
Anatomisches Institut, Eberhard-Karls-Universität Tübingen, Germany.
J Comp Physiol A. 1999 Apr;184(4):361-9. doi: 10.1007/s003590050335.
The focal length of the vertebrate eye is a function of wavelength, i.e. the eye suffers from longitudinal chromatic aberration. Chromatic defocus is a particularly severe problem in eyes with high light-gathering ability, since depth of field is small due to a pupillary opening that is large in relation to the focal length of the eye. Calculations show that in such eyes only a narrow spectral band of light can be in focus on the retina. For the major part of the visual spectrum, spatial resolution should be limited by the optics of the eye and far lower than the resolving power achievable by the retinal cone photoreceptor mosaic. To solve this problem, fishes with irises unresponsive to light have developed lenses with multiple focal lengths. Well-focused images are created at the wavelengths of maximum absorbance of all spectral cone types. Multifocal lenses also appear to be present in some terrestrial species. In eyes with mobile irises, multifocal lenses are correlated with pupil shapes that allow all zones of the lens, with different refractive powers, to participate in the imaging process, irrespective of the state of pupil constriction.
脊椎动物眼睛的焦距是波长的函数,即眼睛存在纵向色差。在聚光能力强的眼睛中,色差散焦是一个特别严重的问题,因为相对于眼睛的焦距而言,瞳孔开口较大,导致景深较小。计算表明,在这样的眼睛中,只有很窄的光谱带的光能够聚焦在视网膜上。对于大部分可见光谱来说,空间分辨率应该受眼睛光学系统的限制,远低于视网膜视锥光感受器镶嵌结构所能达到的分辨能力。为了解决这个问题,虹膜对光无反应的鱼类进化出了具有多个焦距的晶状体。在所有光谱视锥类型的最大吸收波长处能形成聚焦良好的图像。多焦点晶状体似乎在一些陆生物种中也存在。在虹膜可活动的眼睛中,多焦点晶状体与瞳孔形状相关,这种瞳孔形状能使具有不同屈光力的晶状体的所有区域都参与成像过程,而与瞳孔收缩状态无关。