Farrell Una C, Briggs Derek E G
Department of Geology and Geophysics, Yale University, PO Box 208109, New Haven, CT 06520-8109, USA.
Proc Biol Sci. 2007 Feb 22;274(1609):499-504. doi: 10.1098/rspb.2006.0063.
A polychaete from the Middle Devonian Arkona Shale at Hungry Hollow, Arkona, Ontario is preserved in three dimensions in pyrite. The prostomium bears a single median antenna, a pair of lateral antennae and a pair of ventral palps. It is assumed to be fused to a reduced peristomium. The anteriormost three pairs of trunk appendages are modified as tentacular cirri, the third long and biramous. The remainder of the finely annulated trunk bears at least 21 similar biramous parapodia, some of which preserve evidence of chaetae. The postsegmental pygidium is very small and may bear up to two pairs of cirri. The polychaete, Arkonips topororum, falls within the Palpata, Aciculata, among the crown group Phyllodocida. Its remarkable preservation highlights the potential of the Arkona Shale to yield other examples of soft-tissue preservation.
在安大略省阿尔科纳的饥饿谷,一块来自中泥盆世阿尔科纳页岩的多毛类动物以三维形态保存在黄铁矿中。口前叶有一根单一的中触角、一对侧触角和一对腹须。据推测,它与一个退化的围口节融合在一起。最前面的三对躯干附肢演变成触手状触须,第三对长且呈双分支状。其余有精细环纹的躯干至少有21对类似的双分支疣足,其中一些保留了刚毛的痕迹。分节后的尾节非常小,可能有多达两对触须。这种多毛类动物,阿尔科纳托波罗多毛虫,属于触须亚纲、针虫目,在叶须虫目冠群之中。它保存得非常完好,凸显了阿尔科纳页岩产出其他软组织保存实例的潜力。