Nishida Harufumi, Pigg Kathleen B, Kudo Kensuke, Rigby John F
Faculty of Science and Engineering, Chuo University, 1-13-27 Kasuga, Bunkyo, Tokyo, 112-8551, Japan.
J Plant Res. 2007 Jul;120(4):539-49. doi: 10.1007/s10265-007-0093-0. Epub 2007 May 30.
This study describes Homevaleia gouldii H. Nishida, Pigg, Kudo et Rigby gen. et sp. nov., an ovule-bearing glossopterid organ, based on a combination of recently collected permineralized specimens from the Late Permian Homevale Station locality in the Bowen Basin of Queensland, Australia, and on previously studied material from the 1977 Gould and Delevoryas study. Homevaleia, which resembles the compression-impression genus Dictyopteridium, is an inrolled megasporophyll with a distinct keel that bears numerous (over 70) stalked ovules on its adaxial surface. Ovules are small, oval, with an elaborate mesh-like structure that is developed from the outermost integumentary layers. Specimens interpreted as representing different developmental stages show there is an apparent interrelationship between megagametophyte development and the opening of the surrounding fertile structure for pollination. Together, new information provided by this material enables better understanding of glossopterid reproductive structure and its function in one distinctive form.
本研究基于近期从澳大利亚昆士兰州博文盆地二叠纪晚期霍姆韦尔站产地采集的矿化标本以及1977年古尔德和德莱沃里亚斯研究中先前已研究的材料，描述了具胚珠舌羊齿类器官新属新种——古尔德霍姆韦尔羊齿（Homevaleia gouldii H. Nishida, Pigg, Kudo et Rigby gen. et sp. nov.）。霍姆韦尔羊齿类似于压型化石属的网叶羊齿（Dictyopteridium），是一种内卷的大孢子叶，具明显的龙骨状突起，其近轴面着生众多（超过70个）具柄胚珠。胚珠小，呈椭圆形，具由最外层珠被层发育而来的精细网状结构。被解释为代表不同发育阶段的标本显示，大配子体发育与周围可育结构开口进行授粉之间存在明显的相互关系。总之，这些材料提供的新信息有助于更好地理解舌羊齿类的生殖结构及其在一种独特形态中的功能。
