Science. 1965 Jul 23;149(3682):426-7. doi: 10.1126/science.149.3682.426.
Polygenic mutations were induced in an inbred strain of Tribolium confusum by exposure to 500 roentgens of gamma radiation. Female progeny of irradiated males bearing induced mutations in the heterozygous state produced significantly more viable offspring than control female progeny of nonirradiated males.
多基因突变是通过将内布拉斯加州的赤拟谷盗暴露在 500 伦琴的γ射线下而诱发的。携带杂合状态下诱导突变的雄性后代的雌性后代比未受照射的雄性后代的雌性后代产生的存活后代明显更多。