Science. 1975 Feb 7;187(4175):445-7. doi: 10.1126/science.187.4175.445.
Comparative study of pollen of the ranalean complex has revealed a remarkable, hitherto unrecognized characteristic of primitive angiosperm pollen, namely, its complete lack of columellae. Pollen with such exine has been desnated atectate anid taxa in the Magnoliaceae. Degenzeriaceae, Eupomatiaceae, Annonraceae, and possibly Himantandraceae and Nymphaeaceae have pollen which is considered to be primitively columellaless.
对蝾螈类复合体的花粉进行比较研究揭示了一个显著的、迄今尚未被认识到的原始被子植物花粉特征，即完全缺乏柱状体。具有这种外壁的花粉被指定为木兰科、八角科、樟科、番荔枝科、可能还有肉豆蔻科和睡莲科的特化类型。
