Réblová Martina
Department of Plant Taxonomy & Biosystematics, Institute of Botany, Academy of Science, Průhonice 252 43, Czech Republic.
Mycologia. 2008 Nov-Dec;100(6):893-901. doi: 10.3852/08-068.
The phylogenetic analyses of partial nucLSU rDNA sequence data of three Jobellisia species indicate that J. rhynchostoma is distinct from the core species of Jobellisia. Jobellisia luteola, the type species of the genus, and J. fraterna reside as a strongly supported monophyletic clade in a basal position in a grouping containing the Diaporthales, the Calosphaeriales and the Togniniaceae, while all phylogenies confirm the placement of J. rhynchostoma within the Sordariales. The new family Jobellisiaceae (incertae sedis) is described for Jobellisia. A new perithecial ascomycete genus, Bellojisia (Lasiosphaeriaceae, Sordariales), is introduced for J. rhynchostoma. The fungus produces nonstromatic, long-necked perithecia with a superficial to semi-immersed pyriform venter and carbonaceous three-layered perithecial wall, 1-septate, hyaline, later brown, reniform to navicular ascospores with a polar germ pore formed in unitunicate asci. The fungus was not observed to produce a conidial anamorph in vitro. Both morphological and molecular data suggest Corylomyces selenosporus of the Sordariales is the closest relative of J. rhynchostoma. The other relatives of Bellojisia (viz. Cercophora, Lasiosphaeria and Podospora) recruit from the Lasiosphaeriaceae (Sordariales). Cercophora and Podospora are shown as polyphyletic within the Sordariales, which is in agreement with previous molecular studies.
对三种约贝氏菌属物种的部分核糖体大亚基（nucLSU）rDNA序列数据进行的系统发育分析表明，喙口约贝氏菌与约贝氏菌属的核心物种不同。该属的模式种黄褐约贝氏菌和兄弟约贝氏菌在包含座囊菌目、球壳目和托格尼亚科的一个类群的基部位置，作为一个得到有力支持的单系分支存在，而所有系统发育分析都证实喙口约贝氏菌位于粪壳菌目内。为约贝氏菌属描述了新科约贝氏菌科（地位未定）。为喙口约贝氏菌引入了一个新的子囊菌有性子囊壳属，即贝洛约贝氏菌属（粪壳菌科，粪壳菌目）。该真菌产生无子座、长颈子囊壳，具表面至半埋生的梨形腹部，以及含碳的三层子囊壳壁，子囊孢子1分隔、透明、后期褐色、肾形至舟形，在单囊壁子囊中形成具极生芽孔。在体外未观察到该真菌产生分生孢子无性型。形态学和分子数据均表明，粪壳菌目中的硒孢棒囊腔菌是喙口约贝氏菌最近的亲缘种。贝洛约贝氏菌属的其他亲缘属（即尾孢壳属、毛球壳属和柄孢壳属）来自毛球壳科（粪壳菌目）。尾孢壳属和柄孢壳属在粪壳菌目中显示为多系的情况，这与之前的分子研究结果一致。
