Center for Theoretical Study, Charles University, Jilská 1, 110 00-CZ Praha 1, Czech Republic Department of Ecology, Faculty of Sciences, Viničná 7, 128 44-CZ Praha 2, Czech Republic Biodiversity and Macroecology Group, Department of Animal & Plant Sciences, University of Sheffield, Sheffield S10 2TN, UK.
Ecol Lett. 2005 May;8(5):487-92. doi: 10.1111/j.1461-0248.2005.00740.x.
Area and available energy are major determinants of species richness. Although scale dependency of the relationship between energy availability and species richness (the species-energy relationship) has been documented, the exact relationship between the species-area and the species-energy relationship has not been studied explicitly. Here we show, using two extensive data sets on avian distributions in different biogeographic regions, that there is a negative interaction between energy availability and area in their effect on species richness. The slope of the species-area relationship is lower in areas with higher levels of available energy, and the slope of the species-energy relationship is lower for larger areas. This three-dimensional species-area-energy relationship can be understood in terms of probabilistic processes affecting the proportions of sites occupied by individual species. According to this theory, high environmental energy elevates species' occupancies, which depress the slope of the species-area curve.
面积和可用能量是决定物种丰富度的主要因素。尽管已有文献记录了能量可利用性与物种丰富度之间的关系（物种-能量关系）存在尺度依赖性，但物种-面积关系和物种-能量关系之间的确切关系尚未被明确研究。在这里，我们使用两个关于不同生物地理区域鸟类分布的广泛数据集表明，在影响物种丰富度方面，可用能量和面积之间存在负相互作用。在可用能量水平较高的地区，物种-面积关系的斜率较低，而对于较大的地区，物种-能量关系的斜率也较低。这种三维的物种-面积-能量关系可以用影响单个物种占据的站点比例的概率过程来理解。根据这一理论，较高的环境能量会提高物种的占有率，从而降低物种-面积曲线的斜率。
