Carrillo-Ruiz H, Morón M A
Escuela de Biología, Benemérita Univ Autónoma de Puebla, México.
Neotrop Entomol. 2011 Mar-Apr;40(2):217-21.
Results of phylogenetic analysis based on 34 morphological characters of 24 species of 11 genera of Hopliinae from Europe, Japan, South Africa, Madagascar, North and Central America, indicates that the genus Hoplia is a monophyletic group with species distributed in Europe, Japan and America. Based in this analysis the Asiatic genus Ectinohoplia is the closest relative of the genus Hoplia, and the South American genus Barybas (Melolonthinae: Macrodactylini) is the sister group of Hopliinae.
基于欧洲、日本、南非、马达加斯加、北美洲和中美洲11个属24个物种的34个形态特征进行的系统发育分析结果表明，Hoplia属是一个单系类群，其物种分布于欧洲、日本和美洲。基于该分析，亚洲的Ectinohoplia属是Hoplia属的近亲，而南美洲的Barybas属（鳃金龟科：大爪鳃金龟族）是Hopliinae亚科的姐妹群。
