Vuco J A, Anastasijević R J
Institut za medicinska istrazivanja, Beograd.
Arh Hig Rada Toksikol. 1990 Mar;41(1):47-56.
Apart from the direct influences of muscle spindle sensory endings on skeletomotor neurones a great deal of their reflex drive is provided indirectly via the gamma-loop. Fusimotor neurones modulate their discharge rate in parallel with that of the skeletomotor neurones during the rising phase, as well as during the maintained reflex tension in m.triceps surae of decerebrated cat, when small tension oscillations are present in the muscle. However, under other conditions of spinal reflex activities either skeletomotor or fusimotor neurones could be activated independently by impulses from muscle receptors. On the ground of the presented results, it can be supposed that fusimotor firing, together with the afferent firing from muscle spindles, does not provide the main stimulus for the reflex activity of skeletomotor neurones but serves as an indicator of the deviations from the expected or planned movement representing, in this way, its model.
除了肌梭感觉末梢对骨骼肌运动神经元的直接影响外,它们的大量反射驱动是通过γ环路间接提供的。在去大脑猫的比目鱼肌上升期以及维持反射张力期间,当肌肉中存在小的张力振荡时,梭内运动神经元与骨骼肌运动神经元的放电频率平行调节。然而,在脊髓反射活动的其他条件下,骨骼肌运动神经元或梭内运动神经元可能被来自肌肉感受器的冲动独立激活。根据所呈现的结果,可以推测梭内运动神经元的放电,与来自肌梭的传入放电一起,并不是骨骼肌运动神经元反射活动的主要刺激因素,而是作为代表预期或计划运动偏差的指标,从而构成其模型。