Likhitrakarn Natdanai, Golovatch Sergei I, Panha Somsak
Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand.
Zookeys. 2011(131):1-161. doi: 10.3897/zookeys.131.1921. Epub 2011 Sep 29.
The large genus Orthomorpha is rediagnosed and is shown to currently comprise 51 identifiable species ranging from northern Myanmar and Thailand in the Northwest to Lombok Island, Indonesia in the Southeast. Of them, 20 species have been revised and/or abundantly illustrated, based on a restudy of mostly type material; further 12 species are described as new: Orthomorpha atypicasp. n., Orthomorpha communissp. n., Orthomorpha isarankuraisp. n., Orthomorpha picturatasp. n., Orthomorpha similanensissp. n., Orthomorpha suberectasp. n., Orthomorpha tuberculiferasp. n.,Orthomorpha subtuberculiferasp. n. and Orthomorpha latitergasp. n., all from Thailand, as well as Orthomorpha elevatasp. n.,Orthomorpha spiniformissp. n. and Orthomorpha subelevatasp. n., from northern Malaysia. The type-species Orthomorpha beaumontii (Le Guillou, 1841) is redescribed in due detail from male material as well, actually being a senior subjective synonym of Orthomorpha spinala (Attems, 1932), syn. n. Two additional new synonymies are proposed: Orthomorpha rotundicollis (Attems, 1937) = Orthomorpha tuberculata (Attems, 1937), syn. n., and Orthomorpha butteli Carl, 1922 = Orthomorpha consocius Chamberlin, 1945, syn. n., the valid names to the left. All species have been keyed and all new and some especially widespread species have been mapped. Further six species, including two revised from type material, are still to be considered dubious, mostly because their paraterga appear to be too narrow to represent Orthomorpha species. A new genus, Orthomorphoidesgen. n., diagnosed versus Orthomorpha through only moderately well developed paraterga, coupled with a poorly bi- or trifid gonopod tip, with at least some of its apical prongs being short spines, is erected for two species: Orthomorpha setosus (Attems, 1937), the type-species, which is also revised from type material, and Orthomorpha exaratus (Attems, 1953), both comb. n. ex Orthomorpha.
大型的正形马陆属被重新界定,目前已知包含51个可识别的物种,分布范围从西北的缅甸北部和泰国至东南的印度尼西亚龙目岛。其中,基于对大部分模式标本的重新研究,20个物种已得到修订和/或有大量插图说明;另有12个物种被描述为新物种:异型正形马陆新种、普通正形马陆新种、伊沙拉库拉正形马陆新种、斑纹正形马陆新种、斯米兰正形马陆新种、近直立正形马陆新种、具瘤正形马陆新种、亚具瘤正形马陆新种和宽背正形马陆新种,均来自泰国,以及高正形马陆新种、棘状正形马陆新种和近高正形马陆新种,来自马来西亚北部。模式种博蒙特正形马陆(Le Guillou,1841)也根据雄性标本进行了详细重新描述,实际上它是棘正形马陆(Attems,1932)的次主观同物异名,新组合。另外还提出了两个新的同物异名:圆颈正形马陆(Attems,1937)=瘤正形马陆(Attems,1937),新组合,以及布特尔正形马陆Carl,1922 = 同群正形马陆Chamberlin,1945,新组合,有效名称在左边。所有物种均已编制检索表,所有新物种以及一些分布特别广泛的物种都已绘制分布图。另外还有6个物种,包括2个根据模式标本修订的物种,仍被视为存疑物种,主要是因为它们的侧背片似乎太窄,无法代表正形马陆属物种。一个新属,拟正形马陆属,通过仅适度发育的侧背片以及发育不良的双叉或三叉生殖肢顶端来与正形马陆属区分,其顶端的一些叉为短刺,该属包含两个物种:模式种刚毛刺正形马陆(Attems,1937),也根据模式标本进行修订,以及裸正形马陆(Attems,1953),均为来自正形马陆属的新组合。