Department of Cell and Structural Biology, University of Manchester, U.K.
Eur J Protistol. 1989 Apr 14;24(3):205-15. doi: 10.1016/S0932-4739(89)80057-4. Epub 2011 Nov 2.
The ciliate Dasytricha ruminantium possesses cilia over the entire surface. Both the oral and somatic kineties are composed of monokinetids comprising a kinetosome, a tangential ribbon of 5 transverse microtubules, a weakly convergent bundle of 5 postciliary microtubules, a short kinetodesmal fibre, and a single microtubule homologous to T(2) of other litostomate ciliates [15]. The reversed orientation of the kineties within and around the vestibulum may be a consequence of the evolutionary migration of the vestibulum from the anterior to the posterior of the cell. The vestibulum leads to the cytostome and a cytopharynx of the rhabdos type [3]. Microtubules lining the exit canals of the posterior contractile vacuole and cytoproct are believed to originate from somatic kinetids. The ecto-endoplasmic boundary layer (eeb) is composed of two microfibrillar layers. A large extension of the eeb connects the vestibulum to the cell cortex but does not form a karyophore as seen in the closely related genus Isotricha [12].
纤毛虫大草履虫体表布满纤毛。口纤毛和体纤毛均由单联体组成，包括基体、5 根横向微管组成的切向带、5 根后纤毛的弱汇聚束、短动基体纤维和与其他缘毛目纤毛虫的 T(2)同源的单根微管[15]。前庭内和周围纤毛的反向排列可能是前庭从细胞的前部向后部进化迁移的结果。前庭通向胞口和原咽[3]。被认为起源于体联体的微管排列在后伸缩泡和胞肛的出口道。外质-内质边界层（eeb）由两个微纤维层组成。eeb 的一个大延伸将前庭与细胞皮层连接起来，但不像在密切相关的属等缘虫中那样形成核质体[12]。
