Kauffman Stuart A
Institute for Systems Biology, Seattle, WA 98109, USA.
Biosystems. 2014 Sep;123:3-8. doi: 10.1016/j.biosystems.2014.03.004. Epub 2014 Apr 3.
Despite Darwin, we remain children of Newton and dream of a grand theory that is epistemologically complete and would allow prediction of the evolution of the biosphere. The main purpose of this article is to show that this dream is false, and bears on studying patterns of evolution. To do so, I must justify the use of the word "function" in biology, when physics has only happenings. The concept of "function" lifts biology irreducibly above physics, for as we shall see, we cannot prestate the ever new biological functions that arise and constitute the very phase space of evolution. Hence, we cannot mathematize the detailed becoming of the biosphere, nor write differential equations for functional variables we do not know ahead of time, nor integrate those equations, so no laws "entail" evolution. The dream of a grand theory fails. In place of entailing laws, I propose a post-entailing law explanatory framework in which Actuals arise in evolution that constitute new boundary conditions that are enabling constraints that create new, typically unprestatable, adjacent possible opportunities for further evolution, in which new Actuals arise, in a persistent becoming. Evolution flows into a typically unprestatable succession of adjacent possibles. Given the concept of function, the concept of functional closure of an organism making a living in its world becomes central. Implications for patterns in evolution include historical reconstruction, and statistical laws such as the distribution of extinction events, or species per genus, and the use of formal cause, not efficient cause, laws.
尽管有达尔文,但我们仍然是牛顿的信徒,梦想着有一个在认识论上完备的宏大理论,它能让我们预测生物圈的进化。本文的主要目的是表明这个梦想是错误的,并且与研究进化模式有关。为此,我必须为生物学中“功能”一词的使用进行辩护,因为在物理学中只有事件。“功能”的概念将生物学不可还原地提升到物理学之上,因为正如我们将看到的,我们无法预先陈述不断出现的新的生物学功能,而这些功能构成了进化的相空间。因此,我们无法将生物圈的详细变化过程数学化,也无法为我们事先不知道的功能变量写出微分方程,更无法对这些方程进行积分,所以不存在“支配”进化的定律。宏大理论的梦想破灭了。取而代之的是一种后支配定律的解释框架,在这个框架中,进化中出现的实际情况构成了新的边界条件,这些条件是促成性约束,为进一步的进化创造了新的、通常无法预先陈述的相邻可能性机会,在这个过程中不断有新的实际情况出现。进化流入一系列通常无法预先陈述的相邻可能性中。鉴于功能的概念,生物体在其生存世界中的功能封闭概念就变得至关重要。对进化模式的影响包括历史重建,以及统计定律,如灭绝事件的分布或每个属的物种数量,还有使用形式因而非动力因的定律。