Merle Didier, Pacaud Jean-Michel, Métais Grégoire, Bartolini Annachiara, Lashari Rafiq A, Brohi Imdad A, Solangi Sarfraz H, Marivaux Laurent, Welcomme Jean-Loup
Sorbonne Universités - CR2P - MNHN, CNRS, UPMC-Paris6. Muséum national d'Histoire naturelle, Département Histoire de la Terre. Case postale 38, 8 rue Buffon, F-75231 Paris cedex 05 (France); Email:
Sorbonne Universités - CR2P - MNHN, CNRS, UPMC-Paris6. Muséum national d'Histoire naturelle, Département Histoire de la Terre. Case postale 38, 8 rue Buffon, F-75231 Paris cedex 05 (France); Email: unknown.
Zootaxa. 2014 Jun 27;3826(1):101-38. doi: 10.11646/zootaxa.3826.1.3.
The paleobiodiversity of the Volutidae (Mollusca: Gastropoda) of the Ranikot Group (Sindh, Pakistan) and particularly of the Lakhra Formation (SBZ 5 biozone, Earliest Eocene), is reconsidered on the basis of new material collected during recent field trips. Ten new species are described (Mitreola brohii sp. nov., Lyrischapa vredenburgi sp. nov., L. brevispira sp. nov., Athleta (Volutopupa) citharopsis sp. nov., A. (Volutocorbis) lasharii sp. nov., Volutilithes welcommei sp. nov., V. sindhiensis sp. nov., Pseudaulicina coxi sp. nov., Sindhiluta lakhraensis sp. nov. and Pakiluta solangii sp. nov.) and one species is in open nomenclature (Lyria sp.). Three new genera are described: Lyriopsis gen. nov. [Volutinae, ?Lyriini, type species: Lyriopsis cossmanni (Vredenburg, 1923)], Sindhiluta gen. nov. [Volutilithinae, type species: Sindhiluta lakhraensis n. sp.] and Pakiluta gen. nov. [?Volutodermatinae, type species: Pakiluta solangii n. sp.]. Two new combinations are proposed: Lyriopsis cossmanni (Vredenburg, 1923) comb. nov. and Athleta (Volutopupa) intercrenatus (Cossmann & Pissarro, 1909) comb. nov. Lectotypes are designated for Lyria cossmanni Vredenburg, 1923, L. feddeni Vredenburg, 1923, Volutospina noetlingi Cossmann & Pissarro, 1909, V. intercrenata Cossmann & Pissarro, 1909 and Athleta (Volutocorbis) victoriae Vredenburg, 1923. With 21 species, this volutid fauna is the most diverse recorded from the Tethys Ocean during Earliest Eocene time. The assemblage is characterized by a strong turnover marked by regional speciation and the appearance of many western Tethyan invaders. Although at the species level, the assemblage documents a strong provincialism, at the genus level, the high number of shared genera between Eastern Tethyan and Old World Tethyan realms begins a phase of long-term homogeneity of volutid assemblages from the Tethyan paleobiogeographic province.
基于近期实地考察收集的新材料，对巴基斯坦信德省拉尼科特群尤其是拉克拉组（始新世早期SBZ 5生物带）的梭螺科（软体动物：腹足纲）古生物多样性进行了重新审视。描述了10个新物种（布氏米锥螺新种Mitreola brohii sp. nov.、弗雷登伯格氏丽螺新种Lyrischapa vredenburgi sp. nov.、短螺肋丽螺新种L. brevispira sp. nov.、西塔螺形 Athleta（Volutopupa）新种A. (Volutopupa) citharopsis sp. nov.、拉沙氏A.（Volutocorbis）新种A. (Volutocorbis) lasharii sp. nov.、韦尔科姆氏旋梯螺新种Volutilithes welcommei sp. nov.、信德旋梯螺新种V. sindhiensis sp. nov.、考氏假管螺新种Pseudaulicina coxi sp. nov.、拉克拉信德螺新种Sindhiluta lakhraensis sp. nov.和索朗吉氏巴基螺新种Pakiluta solangii sp. nov.），还有1个物种为待定名称（丽螺属未定种Lyria sp.）。描述了3个新属：丽螺形属Lyriopsis gen. nov. [梭螺亚科，？丽螺族，模式种：科斯曼丽螺形螺Lyriopsis cossmanni（弗雷登伯格，1923年）]、信德螺属Sindhiluta gen. nov. [旋梯螺亚科，模式种：拉克拉信德螺Sindhiluta lakhraensis新种]和巴基螺属Pakiluta gen. nov. [？管螺亚科，模式种：索朗吉氏巴基螺Pakiluta solangii新种]。提出了2个新组合：科斯曼丽螺形螺Lyriopsis cossmanni（弗雷登伯格，1923年）新组合comb. nov.和间圆 Athleta（Volutopupa）新组合A. (Volutopupa) intercrenatus（科斯曼和皮萨罗，1909年）新组合comb. nov.。为1923年的科斯曼丽螺Lyria cossmanni、1923年的费登氏丽螺L. feddeni、1909年的诺伊特林氏旋梯螺Volutospina noetlingi、1909年的间圆旋梯螺V. intercrenata以及1923年的维多利亚 Athleta（Volutocorbis）指定了选模标本。这个梭螺动物群有21个物种，是始新世早期从特提斯洋记录到的最多样化的动物群。该组合的特征是有明显的更替，表现为区域物种形成以及许多西特提斯洋入侵者的出现。虽然在物种层面，该组合显示出很强的地方特有性，但在属层面，东特提斯洋和旧世界特提斯洋区域之间大量共享的属开启了特提斯洋古生物地理省梭螺组合长期同质化的阶段。
