Lambkin Kevin J
Queensland Museum, South Brisbane, Queensland, Australia 4101.; Email:
Zootaxa. 2015 Mar 20;3936(3):357-74. doi: 10.11646/zootaxa.3936.3.3.
Species of the extinct family Dysmorphoptilidae with distinctly punctate and emarginate tegmina are one of the most characteristic elements of the hemipteran fauna of the three Queensland Triassic fossil insect-bearing formations-the Middle Triassic (Anisian) Gayndah Formation at Gayndah, the Late Triassic (Norian) Mount Crosby Formation at Mount Crosby, and the Late Triassic (Norian) Blackstone Formation at Mount Crosby and Dinmore. Eight species in five genera have been identified: Mesocixius triassicus Tillyard, 1919 (Denmark Hill), Mesocixius parvus (Evans) comb. nov., 1956 (Mount Crosby), Triassocixius australicus Tillyard, 1919 (Denmark Hill), Carsburgia knezouri gen. et sp. nov. (Dinmore), Dysmorphoptiloides elongata Evans, 1956 (Mount Crosby), Dysmorphoptiloides ellisi sp. nov. (Gayndah), Tennentsia princeps sp. nov. (Mount Crosby), and Tennentsia evansi sp. nov. (Gayndah). Synapomorphies are proposed to suggest that Triassocixius, Carsburgia, Dysmorphoptiloides and Tennentsia are monophyletic. Mesocixius may be paraphyletic, its two species sharing only apparently plesiomorphic characters. The limits and relationships of the over 15 described world genera of the Dysmorphoptilidae remain poorly known, but examination of the Queensland species has identified several characters which may be of cladistic value: the presence or absence of a strigil, the degree of tegmen emarginations and form of the resultant apical lobe, the presence and form of swellings on the claval margin, the pattern of tegmen punctation, the relativities of the primary forks of R and RA, the branching pattern of RA1, the branching and orientation of RA2,and the presence or absence of a fusions between RA and RP distally and between M and CuA basally.
已灭绝的畸形翅蝽科物种，其翅鞘具有明显的点状和缺刻，是昆士兰三叠纪三个含化石昆虫地层半翅目动物群最具特色的元素之一——位于盖因达的中三叠世（安尼西阶）盖因达组、位于克罗斯比山的晚三叠世（诺利阶）克罗斯比山组，以及位于克罗斯比山和丁莫尔的晚三叠世（诺利阶）布莱克斯通组。已鉴定出五个属的八个物种：1919年的三叠纪中翅蝽（丹麦山）、1956年的小中翅蝽（埃文斯）新组合、1919年的澳大利亚三叠翅蝽（蒂利亚德）（丹麦山）、新属新种克内佐尔卡氏蝽（丁莫尔）、1956年的细长畸形拟蝽（埃文斯）（克罗斯比山）、新种埃利斯畸形拟蝽（盖因达）、新种王子坦嫩特蝽（克罗斯比山）和新种埃文斯坦嫩特蝽（盖因达）。提出了共同衍征以表明三叠翅蝽属、卡氏蝽属、畸形拟蝽属和坦嫩特蝽属是单系的。中翅蝽属可能是并系的，其两个物种仅共享明显的近祖性状。已描述的世界上超过15个畸形翅蝽科属的界限和关系仍然知之甚少，但对昆士兰物种的研究确定了几个可能具有分支分类价值的特征：是否存在刮器、翅鞘缺刻的程度和由此产生的顶端叶的形状、爪片边缘肿胀的存在和形状、翅鞘点状的模式、R和RA初级分支的相对关系、RA1的分支模式、RA2的分支和方向，以及RA和RP在远端以及M和CuA在基部之间是否融合。
