Lessner Emily J, Stocker Michelle R, Smith Nathan D, Turner Alan H, Irmis Randall B, Nesbitt Sterling J
Department of Geosciences, Virginia Polytechnic Institute and State University (Virginia Tech), Blacksburg, VA, United States; Department of Biological Sciences, Virginia Polytechnic Institute and State University (Virginia Tech), Blacksburg, VA, United States.
Department of Geosciences, Virginia Polytechnic Institute and State University (Virginia Tech) , Blacksburg, VA , United States.
PeerJ. 2016 Sep 6;4:e2336. doi: 10.7717/peerj.2336. eCollection 2016.
Rauisuchids are large (2-6 m in length), carnivorous, and quadrupedal pseudosuchian archosaurs closely related to crocodylomorphs. Though geographically widespread, fossils of this clade are relatively rare in Late Triassic assemblages. The middle Norian (∼212 Ma) Hayden Quarry of northern New Mexico, USA, in the Petrified Forest Member of the Chinle Formation, has yielded isolated postcranial elements and associated skull elements of a new species of rauisuchid. Vivaron haydeni gen. et. sp. nov. is diagnosed by the presence of two posteriorly directed prongs at the posterior end of the maxilla for articulation with the jugal. The holotype maxilla and referred elements are similar to those of the rauisuchid Postosuchus kirkpatricki from the southwestern United States, but V. haydeni shares several maxillary apomorphies (e.g., a distinct dropoff to the antorbital fossa that is not a ridge, a straight ventral margin, and a well defined dental groove) with the rauisuchid Teratosaurus suevicus from the Norian of Germany. Despite their geographic separation, this morphological evidence implies a close phylogenetic relationship between V. haydeni and T. suevicus. The morphology preserved in the new Hayden Quarry rauisuchid V. haydeni supports previously proposed and new synapomorphies for nodes within Rauisuchidae. The discovery of Vivaron haydeni reveals an increased range of morphological disparity for rauisuchids from the low-paleolatitude Chinle Formation and a clear biogeographic connection with high paleolatitude Pangea.
劳氏鳄科动物体型巨大(体长2至6米),是肉食性四足伪鳄类主龙,与鳄形类关系密切。尽管在地理上分布广泛,但该类群的化石在晚三叠世组合中相对稀少。美国新墨西哥州北部中诺利阶(约2.12亿年前)的海登采石场,位于钦尔组的石化森林段,产出了一种新劳氏鳄科动物的孤立颅后骨骼元素和相关头骨元素。海登维瓦龙(Vivaron haydeni)新属新种的特征是,上颌后端有两个向后的尖叉,用于与颧骨铰接。正模上颌骨及参考骨骼元素与美国西南部劳氏鳄科的柯氏后鳄(Postosuchus kirkpatricki)相似,但海登维瓦龙与德国诺利阶的苏氏狂齿鳄(Teratosaurus suevicus)共享几个上颌独征(例如,通向眶前窝的明显落差而非脊状,腹侧边缘笔直,以及齿槽清晰)。尽管它们在地理上相隔甚远,但这种形态学证据表明海登维瓦龙与苏氏狂齿鳄之间存在密切的系统发育关系。新的海登采石场劳氏鳄科动物海登维瓦龙所保存的形态,支持了先前提出的以及劳氏鳄科内各分支的新共源性状。海登维瓦龙的发现揭示了来自低古纬度钦尔组的劳氏鳄科动物在形态差异上的扩大,以及与高古纬度泛大陆的明确生物地理联系。