Kullander Sven O, Norén Michael
Department of Zoology, Swedish Museum of Natural History, PO Box 50007, SE-104 05 Stockholm, Sweden.; Email:
Zootaxa. 2016 Oct 26;4178(4):535-546. doi: 10.11646/zootaxa.4178.4.5.
Danio htamanthinus, new species, is described from small streams in the vicinity of Htamanthi on the middle Chindwin River. It is most similar to D. choprae and D. flagrans from the Ayeyarwaddy River drainage, sharing the same elaborate colour pattern with dark vertical bars and a red interstripe along the posterior side, but is distinguished by the absence of a P+1 stripe and presence of a P stripe represented only by small spots. The uncorrected p-distance in the mitochondrial COI gene separates D. htamanthinus from D. choprae by 4.3% and from D. flagrans by 7.5%. The largest specimen is only 22.9 mm in standard length (male holotype), but the holotype and one other male, 19.5 mm SL, feature sex-specific pectoral-fin tubercles, and a female as small as 16.6 mm SL has ripening ova. In a phylogenetic analysis based on COI sequences, D. htamanthinus is sister taxon of D. flagrans+D. choprae, and those three species are sister group of D. margaritatus+D.erythromicron.
丹尼奥·赫塔曼蒂努斯，新物种，是在钦敦江中游赫塔曼蒂附近的小溪中发现的。它与伊洛瓦底江流域的乔普拉丹尼奥和弗拉格兰斯丹尼奥最为相似，具有相同的精致体色模式，有深色垂直条纹和沿后侧的红色条纹间带，但区别在于没有P + 1条纹，且P条纹仅由小斑点表示。线粒体COI基因中的未校正p距离将赫塔曼蒂努斯丹尼奥与乔普拉丹尼奥分开4.3%，与弗拉格兰斯丹尼奥分开7.5%。最大的标本标准长度仅为22.9毫米（雄性正模标本），但正模标本和另一只19.5毫米标准长度的雄性具有性别特异性的胸鳍瘤，一只仅为16.6毫米标准长度的雌性有成熟的卵子。在基于COI序列的系统发育分析中，赫塔曼蒂努斯丹尼奥是弗拉格兰斯丹尼奥+乔普拉丹尼奥的姐妹分类单元，这三个物种是珍珠丹尼奥+红斑丹尼奥的姐妹群。
