Evolution of cooperative imitators in social networks.

Many evolutionary game models for network reciprocity are based on an imitation dynamics, yet how semirational imitators prevail has seldom been explained. Here we use a model to investigate the coevolutionary dynamics of cooperation and partnership adjustment in a polygenic population of semirational imitators and rational payoff maximizers. A rational individual chooses a strategy best responding to its neighbors when updating strategy and switches to a new partner who can bring it the maximal payoff from all candidates when adjusting the partnership. In contrast, a semirational individual imitates its neighbor's strategy directly and adjusts its partnership based upon a simple reputation rule. Individual-based simulations show that cooperation cannot evolve in a population of all best responders even if they can switch their partners to somebody who can reward them best in game playing. However, when imitators exist, a stable community that consists of cooperative imitators emerges. Further, we show that a birth-death selection mechanism can eliminate all best responders, cultivating a social regime of all cooperative imitators. Compared with parallel simulations that assume fixed networks, cooperative imitators are evolutionarily favored, provided they are able to adjust their partners.