Gittenberger Edmund, Sherub Sherub, Stelbrink Björn
Naturalis Biodiversity Center, P.O. Box 9517, NL-2300RA Leiden, The Netherlends.
Ugyen Wangchuck Institute for Conservation and Environment, Bumthang, Bhutan.
Zookeys. 2017 Jun 8(679):21-28. doi: 10.3897/zookeys.679.13326. eCollection 2017.
The occurrence of at least one species of in Bhutan, viz. , is confirmed by DNA sequencing. A second unnamed species from Bhutan, that might be congeneric, is known from only a single shell. According to the molecular analysis, is most closely related to a still undescribed species from China. These two species together with and , both also from China, form a well supported clade. Awaiting additional molecular data, the apparent inconsistency regarding versus is not dealt with here. The extant true sister species of could be among the four SE Himalayan species from Bhutan and Nepal that are classified with on the basis of conchological data only.
不丹至少有一种[物种名称未给出]的存在通过DNA测序得到了证实。不丹还有第二种未命名的物种，可能是同属的，目前仅知有一个单独的壳。根据分子分析，[物种名称未给出]与中国一种仍未描述的[物种名称未给出]物种关系最为密切。这两个物种与同样来自中国的[物种名称未给出]和[物种名称未给出]一起，形成了一个得到充分支持的进化枝。由于等待更多分子数据，这里暂不处理关于[物种名称未给出]与[物种名称未给出]的明显不一致之处。[物种名称未给出]现存的真正姊妹物种可能是不丹和尼泊尔的四个喜马拉雅东南部物种之一，它们仅根据贝壳学数据被归类为[物种名称未给出]。
