Development of inflorescence architecture is controlled by genetic regulatory networks. (), (), () and () are core regulators for inflorescence development. To understand the regulation of compound inflorescence development, we characterized mutants of corresponding orthologous genes, , , () and , in , and analyzed expression patterns of these genes. Results indicate that , , and play specific roles in identity determination of primary inflorescence meristems, secondary inflorescence meristems, floral meristems and common primordia, respectively. Double mutation of and transforms compound inflorescences to simple flowers, whereas single mutation of changes the inflorescence branching pattern from monopodial to sympodial. Double mutant completely loses floral meristem identity. We conclude that inflorescence architecture in is controlled by spatiotemporal expression of , , and through reciprocal repression. Although this regulatory network shares similarity with the pea model, it has specificity in regulating inflorescence architecture in This study establishes as an excellent genetic model for understanding compound inflorescence development in related legume crops.