Santos-Silva Antonio, Wappes James E, Galileo Maria Helena M
Museu de Zoologia, Universidade de São Paulo, São Paulo, SP, Brazil..
Zootaxa. 2018 Jan 25;4375(4):451-501. doi: 10.11646/zootaxa.4375.4.1.
Estoloides (Parestoloides) Breuning, 1940, and Mimestoloides Breuning, 1974 are synonymized with Estoloides Breuning, 1940. Mimestoloides Breuning, 1980 is synonymized with Spinestoloides Breuning, 1954. Estoloides aurantius Martins Galileo, 2010 is synonymized with E. scabracaulis Chemsak Noguera, 1993. Estoloides modica Chemsak Noguera, 1993 is synonymized with E. chamelae Chemsak Noguera, 1993. Estoloides (Spinestoloides) spinipennis Breuning, 1974, is synonymized with Esthlogena monticola (Fisher, 1942). Estoloides (Spinestoloides) Breuning, 1954 is considered a genus different from Estoloides. Notes on Pseudestoloides Breuning and Heyrovsky, 1961, Pseudestoloides costaricensis Breuning and Heyrovsky, 1961, Pseudestoloides affinis Martins and Galileo, 2009, and on the type of Estoloides (Parestoloides) costaricensis Breuning, 1940, are provided. Pseudestoloides hiekei Breuning, 1974, is transferred to Estoloides. The following new combinations are established: Spinestoloides monticola; Spinestoloides fasciatus (Martins Galileo, 2010); Spinestoloides benardi (Breuning, 1980); Estoloides andresi (Breuning, 1974); and Estoloides hiekei (Breuning, 1974). With the synonymy of Estoloides (Parestoloides), the following species have the subgenus removed from their names: Estoloides basigranulata Breuning, 1943: E. costaricensis Breuning, 1940; E. esthlogenoides Breuning, 1943; and E. parva Breuning, 1940. The following new species are described: Estoloides vandenberghei, from Nicaragua; E. maesi, from Nicaragua; E. nayeliae, from Mexico; E. noguerai, from Mexico and Nicaragua; E. chiapensis, from Mexico; E. pubescens, from Mexico; E. schusteri, from Guatemala; E. dthomasi, from Mexico; E. morrisi, from Mexico; E. giesberti, from Honduras; E. uyucana, from Honduras; Spinestoloides hefferni from Panama; and Pseudestoloides bingkirki from Nicaragua. The male of Spinestoloides spinipennis (Breuning, 1954), and the female of Estoloides prolongata are described for the first time. Keys to separate Estoloides, Pseudestoloides and Spinestoloides, from each other, and to species of Pseudestoloides and Spinestoloides are provided. Estoloides chamelae is newly recorded from Chiapas, Colima, Oaxaca and Guerrero in Mexico; E. prolongata is newly recorded from Costa Rica; and E. rufipes is newly recorded from the Brazilian state of Espírito Santo. The holotypes of Estoloides alboscutellaris Breuning, 1943 (Fig. 13), E. annulicornis Breuning, 1940 (Fig. 14), and E. reflexa Breuning, 1940 (Fig. 15) are also figured for the first time.
1940年的埃斯托洛德属(拟埃斯托洛德亚属)布雷宁和1974年的拟埃斯托洛德属布雷宁被同物异名于1940年的埃斯托洛德属布雷宁。1980年的拟埃斯托洛德属布雷宁被同物异名于1954年的刺埃斯托洛德属布雷宁。2010年的橙黄埃斯托洛德马丁斯和伽利略被同物异名于1993年的糙茎埃斯托洛德切姆萨克和诺格拉。1993年的小型埃斯托洛德切姆萨克和诺格拉被同物异名于1993年的查梅拉埃斯托洛德切姆萨克和诺格拉。1974年的(刺埃斯托洛德属)具刺埃斯托洛德布雷宁被同物异名于1942年的山地埃斯托洛根纳(费希尔)。1954年的(刺埃斯托洛德属)埃斯托洛德布雷宁被认为是一个与埃斯托洛德属不同的属。提供了关于1961年的伪埃斯托洛德布雷宁和海罗夫斯基、1961年的哥斯达黎加伪埃斯托洛德布雷宁和海罗夫斯基、2009年的近缘伪埃斯托洛德马丁斯和伽利略以及1940年的(拟埃斯托洛德亚属)哥斯达黎加埃斯托洛德布雷宁模式种的注释。1974年的希克伪埃斯托洛德布雷宁被转移到埃斯托洛德属。建立了以下新组合:山地刺埃斯托洛德;带纹刺埃斯托洛德(马丁斯和伽利略,2010);贝纳尔迪刺埃斯托洛德(布雷宁,1980);安德雷斯埃斯托洛德(布雷宁,1974);以及希克埃斯托洛德(布雷宁,1974)。随着埃斯托洛德属(拟埃斯托洛德亚属)的同物异名,以下物种的亚属从其名称中去除:1943年的具基粒埃斯托洛德布雷宁;1940年的哥斯达黎加埃斯托洛德布雷宁;1943年的类埃斯托洛根纳埃斯托洛德布雷宁;以及1940年的小型埃斯托洛德布雷宁。描述了以下新物种:来自尼加拉瓜的范登贝赫埃斯托洛德;来自尼加拉瓜的梅西埃斯托洛德;来自墨西哥的纳耶利亚埃斯托洛德;来自墨西哥和尼加拉瓜的诺格拉埃斯托洛德;来自墨西哥的恰帕斯埃斯托洛德;来自墨西哥的柔毛埃斯托洛德;来自危地马拉的舒斯特埃斯托洛德;来自墨西哥的托马斯埃斯托洛德;来自墨西哥的莫里斯埃斯托洛德;来自洪都拉斯的吉斯贝蒂埃斯托洛德;来自洪都拉斯的尤卡纳埃斯托洛德;来自巴拿马的赫弗尼刺埃斯托洛德;以及来自尼加拉瓜的宾基尔基伪埃斯托洛德。首次描述了1954年的具刺刺埃斯托洛德(布雷宁)的雄性和延长埃斯托洛德雌虫。提供了区分埃斯托洛德属、伪埃斯托洛德属和刺埃斯托洛德属以及伪埃斯托洛德属和刺埃斯托洛德属物种的检索表。查梅拉埃斯托洛德在墨西哥的恰帕斯、科利马、瓦哈卡和格雷罗新有记录;延长埃斯托洛德在哥斯达黎加新有记录;以及红足埃斯托洛德在巴西的圣埃斯皮里图州新有记录。1943年的白盾埃斯托洛德布雷宁(图13)、1940年的环角埃斯托洛德布雷宁(图14)和1940年的反曲埃斯托洛德布雷宁(图15)的正模标本也首次绘图。