Couri MÁrcia S, Pont Adrian C
Museu Nacional, Quinta da Boa Vista, São Cristóvão, Rio de Janeiro, 20.940-040, RJ, Brazil..
Zootaxa. 2018 Mar 21;4399(2):233-247. doi: 10.11646/zootaxa.4399.2.7.
The morphology of the male terminalia of fourteen African species of Helina Robineau-Desvoidy, 1830 (Diptera, Muscidae) is described and illustrated: H. dorsalis (Stein, 1914); H. emdeni Pont, 1980, H. fuscibasis Emden, 1951; H. gracilior Emden, 1951; H. hirtipes metatarsalis Emden, 1951, H. juxtamedialis Emden, 1951; H. lasiopa Emden, 1951; H. mollis (Stein, 1906); H. naivashensis Emden, 1951, stat. nov. (herein raised to species rank); H. nemoralis (Stein, 1913); H. novarae (Schiner, 1868), H. penicillata Emden, 1951; H. quadruplex (Stein, 1913); and H. trinubilifera (Malloch, 1921). These species demonstrate that the terminalia can be very varied, with different shapes of sternite 5, sometimes with very strong and long setae, and a short or elongated cercal plate and surstylus that sometimes can bear spines. Some of these species also share other external characters that are unusual among Helina, such as the arrangement of the katepisternal setae as an equilateral triangle, the absence of an anterodorsal seta on mid tibia, and wings with dark clouds. Most of these species are well described in the literature, but the male terminalia have never been studied in detail. The material studied here is deposited in the Natural History Museum (BMNH), London, United Kingdom.
描述并图示了1830年命名的非洲14种黑蝇属（Helina Robineau-Desvoidy，双翅目，蝇科）雄性尾器的形态：背黑蝇（Helina dorsalis (Stein, 1914)）；埃姆登黑蝇（Helina emdeni Pont, 1980），褐基黑蝇（Helina fuscibasis Emden, 1951）；纤小黑蝇（Helina gracilior Emden, 1951）；毛跗黑蝇（Helina hirtipes metatarsalis Emden, 1951），近中黑蝇（Helina juxtamedialis Emden, 1951）；毛鬃黑蝇（Helina lasiopa Emden, 1951）；柔滑黑蝇（Helina mollis (Stein, 1906)）；奈瓦沙黑蝇（Helina naivashensis Emden, 1951，新地位（在此提升为物种等级））；林栖黑蝇（Helina nemoralis (Stein, 1913)）；诺瓦拉黑蝇（Helina novarae (Schiner, 1868)），毛状黑蝇（Helina penicillata Emden, 1951）；四鬃黑蝇（Helina quadruplex (Stein, 1913)）；以及三云黑蝇（Helina trinubilifera (Malloch, 1921)）。这些物种表明，尾器可能非常多样，第5腹板形状各异，有时有非常强壮且长的刚毛，尾铗板和阳茎侧突短或细长，有时可带有刺。其中一些物种还具有黑蝇属中不常见的其他外部特征，如中侧片刚毛排列成等边三角形、中胫无前背刚毛以及翅膀有乌云状斑纹。这些物种中的大多数在文献中有很好的描述，但雄性尾器从未被详细研究过。此处研究的材料存于英国伦敦的自然历史博物馆（BMNH）。
