Kammerer Christian F, Masyutin Vladimir
North Carolina Museum of Natural Sciences, Raleigh, NC, USA.
Vyatka Paleontological Museum, Kirov, Russia.
PeerJ. 2018 Jun 8;6:e4933. doi: 10.7717/peerj.4933. eCollection 2018.
A new therocephalian taxon ( gen. et sp. nov.) is described based on a nearly complete skull and partial postcranium from the Permian Kotelnich locality of Russia. displays an unusual mixture of primitive ("pristerosaurian") and derived (eutherocephalian) characters. Primitive features of include extensive dentition on the palatal boss and transverse process of the pterygoid, paired vomers, and a prominent dentary angle; derived features include the absence of the postfrontal. can be distinguished from all other therocephalians by its autapomorphic dental morphology, with roughly denticulated incisors and postcanines. Phylogenetic analysis recovers as a non-lycosuchid, non-scylacosaurid therocephalian situated as sister-taxon to Eutherocephalia. The identification of as the largest predator from Kotelnich indicates that therocephalians acted as apex predators in middle-late Permian transition ecosystems in Russia, corroborating a pattern observed in South African faunas. However, other aspects of the Kotelnich fauna, and Permian Russian tetrapod faunas in general, differ markedly from those of South Africa and suggest that Karoo faunas are not necessarily representative of global patterns.
基于来自俄罗斯二叠纪科捷尔尼奇地区的一个近乎完整的头骨和部分颅后骨骼，描述了一个新的兽头类分类单元（新属及新种）。它展现出原始（“前龙类”）和衍生（真兽头类）特征的不寻常混合。其原始特征包括翼骨的腭突和横突上有广泛的齿列、成对的犁骨以及突出的齿骨角；衍生特征包括没有额后骨。它可以通过其自近裔性状的牙齿形态与所有其他兽头类区分开来，其门齿和犬后齿大致有细齿。系统发育分析将其恢复为一个非狼蜥兽类、非锯颌兽类的兽头类，作为真兽头类的姐妹分类单元。将其鉴定为科捷尔尼奇最大的捕食者表明，兽头类在俄罗斯中二叠世过渡生态系统中充当顶级捕食者，这证实了在南非动物群中观察到的一种模式。然而，科捷尔尼奇动物群的其他方面，以及一般的俄罗斯二叠纪四足动物群，与南非的动物群有显著差异，这表明卡鲁动物群不一定代表全球模式。
