Kulikovskiy M S, Andreevab S A, Guseva E S, Kuznetsova I V, Annenkova N V
Izv Akad Nauk Ser Biol. 2016 Sep(5):468-478.
The phylogenetic position of the genera Rossithidium and Psammothidium was studied using the plastid gene rbcL. The genus Rossithidium is not an independent taxon and should be transferred to Psammothidium. Taxa from Psammothidium genus formed a sister, but independent clade to taxa with Achnanthidium. Our data has shown that the order Achnathales was not the natural taxon and should be reduced, but the three families included in it were natural. Genera like Psammothidium (including Rossithidium) and Lemnicola should be included into the family Achnanthidiaceae. Therefore, raphe does not play an important role in diatom taxonomy and its loss occurred many times during the evolution of diatoms.
利用质体基因rbcL研究了Rossithidium属和Psammothidium属的系统发育位置。Rossithidium属不是一个独立的分类单元，应归入Psammothidium属。来自Psammothidium属的分类群形成了一个姐妹群，但与Achnanthidium属的分类群是独立的分支。我们的数据表明，Achnathales目不是一个自然分类单元，应该被缩减，但其中包含的三个科是自然的。像Psammothidium属（包括Rossithidium属）和Lemnicola属应该归入Achnanthidiaceae科。因此，脊在硅藻分类学中并不起重要作用，并且在硅藻进化过程中多次发生丢失。
