Nakano Takafumi, Tomikawa Ko, Grygier Mark J
Department of Zoology, Graduate School of Science, Kyoto University, Kitashirakawa-oiwakecho, Sakyo, Kyoto 606-8502, Japan Department of Science Education, Graduate School of Education, Hiroshima University, 1-1-1 Kagamiyama, Higashihiroshima 739-8524, Japan..
Zootaxa. 2018 Dec 16;4532(1):86-94. doi: 10.11646/zootaxa.4532.1.4.
Two missing syntypes of the Japanese subterranean amphipod Procrangonyx japonicus (Uéno, 1930), the type species of Procrangonyx Schellenberg, 1934, were rediscovered in the collections of the Kyoto University Museum. The morphology of uropod 3, which has been considered the principal diagnostic character of the genus, is redescribed on the basis of one of the syntypes, and the nomenclatural history of the generic names Procrangonyx and Eocrangonyx Schellenberg, 1937 (corrected from 1936) for some Far-Eastern subterranean amphipod species is reviewed. Owing to confusion between the terms "type fixation" and "type designation"-the latter being just one means of accomplishing the former-the view that Procrangonyx is unavailable and invalid has prevailed in recent literature. Procrangonyx was indeed proposed after 1930 with no type species "designation", but under Articles 67.2.1 and 68.3 of the International Code of Zoological Nomenclature, Eucrangonyx japonicus Uéno, 1930 was "fixed" as its type species by monotypy in the original publication. Since a diagnosis of the genus was also provided in the same work, Procrangonyx is available under Article 13.3 of the Code. However, because endopodal segmentation of uropod 3 proves to be variable in P. japonicus, doubt is thrown on the taxonomic distinctness of Procrangonyx vis à vis Pseudocrangonyx Akatsuka Komai, 1922. Additionally, the publication dates of Allocrangonyx Schellenberg, 1937 and Niphargus foreli speziae Schellenberg, 1937 are corrected from 1936.
日本地下双足虾类日本原长额虾(Uéno,1930)的两个缺失的合模式标本在京都大学博物馆的馆藏中被重新发现,该物种是1934年Schellenberg的原长额虾属的模式种。基于其中一个合模式标本,重新描述了第三尾肢的形态,该形态一直被视为该属的主要诊断特征,并回顾了1937年(从1936年修正)Schellenberg为一些远东地下双足虾类物种所命名的原长额虾属和始原长额虾属的命名历史。由于“模式确立”和“模式指定”这两个术语之间的混淆——后者只是实现前者的一种手段——原长额虾属不可用且无效的观点在最近的文献中盛行。原长额虾属确实是在1930年之后提出的,没有“指定”模式种,但根据《国际动物命名法规》第67.2.1条和第68.3条,1930年的日本真长额虾在原始出版物中通过单型性被“确立”为其模式种。由于同一著作中也提供了该属的诊断,根据法规第13.3条,原长额虾属是可用的。然而,由于在日本原长额虾中第三尾肢内肢节段被证明是可变的,因此对原长额虾属相对于1922年Akatsuka Komai的伪原长额虾属的分类独特性产生了怀疑。此外,1937年的异原长额虾属和1937年的福尔氏尼氏真虾的出版日期从1936年修正。