Dalla Vecchia Fabio Marco
Research Group of Mesozoic Faunas, Institut Català de Paleontologia Miquel Crusafont (ICP), Sabadell, Catalonia, Spain.
Museo Friulano di Storia Naturale, Udine, Italy.
PeerJ. 2019 Jul 25;7:e7363. doi: 10.7717/peerj.7363. eCollection 2019.
A new non-monofenestratan pterosaur with multicusped dentition, , is described from the Upper Triassic (middle-upper Norian) of the Carnian Prealps (northeastern Italy). The holotype of preserves a complete mandibular and maxillary dentition, along with a nearly complete premaxillary one, showing unique features. Furthermore, the arrangement of the premaxillary teeth and the shape of jugal, pterygoid, ectopterygoid, scapula and pteroid are unique within non-monofenestratan pterosaurs. is similar and closely related to and , which are also from the Alpine middle-upper Norian of Italy and Austria, respectively. In a parsimony-based phylogenetic analysis, is found to nest within a clade of Triassic pterosaurs composed of , and a trichotomy of , and MCSNB 8950. This unnamed clade is basal within the Pterosauria, but is not the basalmost clade. lies outside this clade and is more derived, making the Eudimorphodontidae paraphyletic. increases the diversity of Triassic pterosaurs and brings the number of pterosaur genera and species in the Dolomia di Forni Formation to four.
一种具有多尖齿列的新型非单窗孔翼龙被描述于意大利东北部卡尼阶前阿尔卑斯地区的上三叠统（诺利阶中上部）。该翼龙的正模标本保存了完整的下颌和上颌齿列，以及近乎完整的前颌齿列，展现出独特的特征。此外，前颌齿的排列以及颧骨、翼骨、外翼骨、肩胛骨和翅骨的形状在非单窗孔翼龙中是独一无二的。它与同样分别来自意大利和奥地利阿尔卑斯地区诺利阶中上部的[相关翼龙名称未给出]和[相关翼龙名称未给出]相似且关系密切。在基于简约法的系统发育分析中，发现它嵌套在一个由[相关翼龙名称未给出]、[相关翼龙名称未给出]以及[相关翼龙名称未给出]、[相关翼龙名称未给出]和MCSNB 8950构成的三分法组成的三叠纪翼龙分支内。这个未命名的分支在翼龙目中处于基部位置，但并非最基部的分支。[相关翼龙名称未给出]位于这个分支之外且更为衍化，这使得双形齿翼龙科成为并系群。它增加了三叠纪翼龙的多样性，并使福尔尼组的翼龙属和种的数量达到四个。
