Sutcharit Chirasak, Naggs Fred, Ablett Jonathan, Sang Pham Van, Panha Somsak
Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand Chulalongkorn University Bangkok Thailand.
Mollusca Section, Invertebrates Division, Department of Life Sciences, The Natural History Museums, London SW7 5BD, UK The Natural History Museums London United Kingdom.
Zookeys. 2019 Nov 4;885:1-14. doi: 10.3897/zookeys.885.38980. eCollection 2019.
Since the time of the original description there have been no precise locality records in Cambodia of (Reeve, 1860) and it was believed to be extinct. In 2012, a joint Natural History Museum survey with Vietnamese colleagues rediscovered living populations of this huge sinistral helicoid snail in a protected area of southern Vietnam. The genitalia and radula morphology are re-assessed and type specimens of all recognised congeners are figured herein. The unique morphological characters of this species are a small and simple penis, well-developed amatorial organ complex that incorporates four amatorial organ ducts, a short gametolytic organ complex and spiked papilla, and radula morphology with unicuspid teeth. The type locality of , which has been contentious, is determined here to be in the vicinity of 'Brelum', Vietnam, near the border with Cambodia. In addition, the nucleotide sequences of barcoding genes COI, 16SrRNA and 28S fragments were provided for further comparison.
自最初描述该物种以来，柬埔寨境内一直没有关于（里夫，1860年）的确切产地记录，人们认为它已经灭绝。2012年，自然历史博物馆与越南同事联合进行的一项调查在越南南部的一个保护区重新发现了这种巨大的左旋螺旋蜗牛的现存种群。本文重新评估了其生殖器和齿舌形态，并绘制了所有公认同属物种的模式标本。该物种独特的形态特征包括一个小而简单的阴茎、发育良好的求偶器官复合体（包含四条求偶器官导管）、一个短的配子溶解器官复合体和带刺乳头，以及具有单尖齿的齿舌形态。一直存在争议的该物种的模式产地在此确定为越南靠近柬埔寨边境的“布雷卢姆”附近。此外，还提供了条形码基因COI、16SrRNA和28S片段的核苷酸序列以供进一步比较。
