Novel mutualists of two species infesting in Indonesia.

Several species in the complex have emerged as important pests of woody plants globally, particularly in habitats where they are invasive aliens. These beetles live in obligate symbioses with fungi in the genus . In this study, we identified spp. and their fungal mutualists that have emerged as pests of planted in Riau, Indonesia. Morphological identification and phylogenetic analyses of the mitochondrial cytochrome oxidase subunit I () gene confirmed that and are the most abundant beetles infesting these trees. Multilocus phylogenetic analyses of their fungal mutualists revealed their nonspecific association with six species. These included and five novel mutualists within the species complex (FSSC), four of which reside in the Ambrosia Clade (AFC). These new species are described here as , and .