Schuh Randall T
Department of Entomology, American Museum of Natural History, New York, New York 10024, U.S.A.
Cladistics. 1991 Jun;7(2):157-189. doi: 10.1111/j.1096-0031.1991.tb00030.x.
A cladistic analysis is performed for 114 species of Pilophorini, using 73 characters with 190 states. Illustrations or figure references to the literature are provided for most characters. The resultant Nelson (strict) consensus cladogram is used as a justification for the recognition of the following genera: Alepidiella Poppius, Aloea Linnavuori, Druthmarus Distant, Hypseloecus Reuter, Neoambonea Schuh, Parambonea Schuh, Parasthenaridea Miller, Pherolepis Kulik (paraphyletic), Pilophorus Hahn, and Sthenaridea Reuter (paraphylectic). Pilophorus indonesicus is proposed as a replacement name for Bilirania sumatrana Schuh (=Pilophorus), a secondary homonym of Pilophorus sumatranus Poppius. A list of currently recognized genera and species with summary distributions and host plant associations, and a key to genera are included. Host associations are plotted on the cladogram to reveal the pattern of host shifts. At the generic level and above, a pattern of colonization, rather than co-evolution, is strongly indicated; at the species level, genera of Pilophorini often show restricted plant-group associations, but no clear pattern of coevolution emerges. Major distributional patterns in the Philophorini are mapped, discussed and compared with historical biogeographic schemes for some other groups. The Pilophorini appear to be of tropical Gondwanan origin with subsequent spread into, and differentiation in, the temperate Northern Hemisphere.
对114种毛角蝽族物种进行了支序分析，使用了73个性状，共190种状态。大多数性状都配有插图或文献中的图注。所得的纳尔逊（严格）合意分支图被用作认可以下属的依据：阿氏角蝽属Poppius、阿洛角蝽属Linnavuori、德鲁氏角蝽属Distant、高氏角蝽属Reuter、新安波角蝽属Schuh、副安波角蝽属Schuh、副强角蝽属Miller、毛鳞角蝽属Kulik（并系）、毛角蝽属Hahn和强角蝽属Reuter（并系）。提出用印度尼西亚毛角蝽取代舒氏苏门答腊双带角蝽（=毛角蝽属），后者是苏门答腊毛角蝽Poppius的次异物同名。文中包含了当前认可的属和物种列表，以及简要分布和寄主植物关联信息，还有一个属的检索表。将寄主关联信息绘制在分支图上以揭示寄主转移模式。在属及以上水平，强烈表明是一种定殖模式，而非协同进化模式；在物种水平，毛角蝽族的属通常显示出对特定植物类群的限制关联，但未出现明显的协同进化模式。文中绘制、讨论并比较了毛角蝽族的主要分布模式与其他一些类群的历史生物地理格局。毛角蝽族似乎起源于热带冈瓦纳地区，随后扩散到北半球温带并在那里分化。
