The spectral sensitivities of the middle- and long-wavelength cones: an extension of the two-colour threshold technique of W S Stiles.

When a tiny centred test flash is presented on a small concentric background, the threshold rises with background radiance more quickly than Weber's law would predict. It is argued that under such conditions it is possible, by means of a test sensitivity method, to isolate either the M-cone or the L-cone types throughout the visible spectrum. As predicted, double-branched M- and L-cone tyr functions are found when the test flash and the field are of the same wavelength. From the independent vertical displacements of the two branches as test wavelength is varied, it is possible to derive spectral sensitivities that agree well with dichromatic sensitivities and König fundamentals. The test sensitivities deviate from pi 4 at longer wavelengths and from pi 5 at shorter wavelengths.