Updates to the carnivore fauna from the Sima de los Huesos.

The Sima de los Huesos site in the Sierra de Atapuerca (Burgos, northern Spain) is a Middle Pleistocene locality with the most important accumulation of humans in the European record for this age. In addition to the hominin collection, the Sima has provided numerous faunal remains. Here, we update the carnivoran mammal fauna from the Sima de los Huesos. The analyzed carnivoran fauna is taxonomically diverse consisting of 11 species from 9 genera and four families. The high taxonomic diversity of carnivorans at the site is interpreted as corresponding to high overall ecosystem productivity. A synthetic description of the carnivoran remains is presented adding new information on the biochronology of the carnivoran species assemblage and its paleoecological association with European Middle Pleistocene hominins. Because of their similarity in taphonomic setting and diets-that the carnivoran fauna and humans were found in the same place and had similar protein-rich diets-clarifying the paleontological and paleoecological context of these carnivorans permits a better understanding of the humans also found at this site.