Bashkuev Alexey S
Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, 117997, Russia. .
Zootaxa. 2022 Dec 23;5222(6):545-577. doi: 10.11646/zootaxa.5222.6.3.
Remarkable new finds of Permotanyderidae, an enigmatic "Diptera-like" mecopteran family hitherto known only from the Upper Permian of Australia, are described from the Upper Permian and transitional Permian-Triassic beds in Russia (northern European Russia, southern Urals, and the Tunguska and Kuznetsk Basins). The new taxa are Permotanyderus sibiricus sp. nov., P. sensibilis sp. nov., P. rakitovi sp. nov., Rudimentanus rieki gen. et sp. nov., R. cisuralicus sp. nov., Permotanyderites pseudopolycentropoides gen. et sp. nov., Chimaeropteron antiquum gen. et sp. nov. and Ch. agmatos sp. nov. The most archaic genus, Chimaeropteron gen. nov., is assigned to the new subfamily Chimaeropterinae. Choristotanyderus nanus Riek, 1953 is transferred to Permotanyderus Riek, 1953. The distinct suctorial mouthparts revealed in P. nanus and the analysis of wing characters provide evidence for the affinity of Permotanyderidae with the long-proboscid families Nedubroviidae and Pseudopolycentropodidae.
在俄罗斯（俄罗斯欧洲北部、南乌拉尔以及通古斯卡和库兹涅茨克盆地）的上二叠统以及二叠纪-三叠纪过渡层中，发现了Permotanyderidae科引人注目的新化石。Permotanyderidae是一个神秘的“类似双翅目”的长翅目昆虫科，此前仅在澳大利亚的上二叠统中被发现。新分类单元包括西伯利亚Permotanyderus sibiricus新种、敏感Permotanyderus sensibilis新种、拉基托夫Permotanyderus rakitovi新种、Rudimentanus rieki新属新种、西西伯利亚Rudimentanus cisuralicus新种、假多中心Permotanyderites pseudopolycentropoides新属新种、古老Chimaeropteron antiquum新属新种以及Chimaeropteron agmatos新种。最古老的属Chimaeropteron新属被归入新亚科Chimaeropterinae。1953年的矮小Choristotanyderus nanus被转移到1953年的Permotanyderus属。在矮小Permotanyderus中发现的独特吸食口器以及对翅特征的分析，为Permotanyderidae科与长喙科Nedubroviidae和Pseudopolycentropodidae的亲缘关系提供了证据。
