Brown John W, Aarvik Leif, Timm Alicia
Department of Entomology; National Museum of Natural History; Washington; DC 20013-7012; USA.
Natural History Museum; University of Oslo; P.O. Box 1172 Blindern NO-0318 Oslo; Norway.
Zootaxa. 2023 Apr 5;5263(2):217-239. doi: 10.11646/zootaxa.5263.2.3.
Gibbalaria Brown and Aarvik, new genus, is described to accommodate the following species: G. divergana Brown and Aarvik, new species (TL: South Africa); G. occidentana Brown and Aarvik, new species (TL: South Africa); G. chiloanei Brown and Aarvik, new species (TL: South Africa); G. scabellana Zeller, 1852, new combination (TL: South Africa); G. mabalingwae Razowski, 2008, status revised, new combination (TL: South Africa); G. longiphallus Brown and Aarvik, new species (TL: South Africa); G. bagamoyo (Razowski & Wojtusiak, 2014), new combination (TL: Tanzania); and G. analcis (Razowski, 2015), new combination (TL: Republic of Cameroon). Gibbalaria sistrata (Meyrick, 1911), new combination (TL: South Africa), is synonymized with G. scabellana, new synonymy. Although the male genitalia of Gibbalaria form two moderately divergent groups, the two groups share extremely similar female genitalia, which feature a broad, somewhat cylindrical antrum, followed immediately anterad by a conspicuously elbowed region of the ductus bursae forming a weak diverticulum on the left side, and a signum in the form of a patch of small arrowhead-shaped sclerites, usually with a shallow pocket. Male genitalia morphology shows many similarities with the Neopotamia group (Olethreutini), Afroploce Aarvik in particular. The monophyly of Gibbalaria is supported by the raised, curled, opalescent scales on the forewing in both sexes, which occurs in a few other genera scattered through the subfamily (e.g., Thylacandra Diakonoff in Grapholitini; Astronauta Diakonoff and Cosmopoda Diakonoff in Olethreutini); the angled diverticulum in the ductus bursae in the female genitalia; the subrectangular flange from the lower margin of the valva immediately basal to the cucullus; and the setose, protruding ridge immediate above the basal cavity, which appears to be homologous with the subbasal process of the valva in many members of the Neopotamia group of genera. In addition, Diakonoffiana graziani Razowski, 2015 (TL: Republic of Cameroon) is transferred to Cosmopoda Diakonoff, resulting in C. graziani (Razowski), new combination.
描述了新属Gibbalaria Brown和Aarvik，包含以下物种：G. divergana Brown和Aarvik，新物种（模式产地：南非）；G. occidentana Brown和Aarvik，新物种（模式产地：南非）；G. chiloanei Brown和Aarvik，新物种（模式产地：南非）；G. scabellana Zeller，1852，新组合（模式产地：南非）；G. mabalingwae Razowski，2008，状态修订，新组合（模式产地：南非）；G. longiphallus Brown和Aarvik，新物种（模式产地：南非）；G. bagamoyo（Razowski & Wojtusiak，2014），新组合（模式产地：坦桑尼亚）；以及G. analcis（Razowski，2015），新组合（模式产地：喀麦隆共和国）。Gibbalaria sistrata（Meyrick，1911），新组合（模式产地：南非），被定为G. scabellana的同物异名，新异名。尽管Gibbalaria的雄性生殖器形成两个适度不同的类群，但这两个类群的雌性生殖器极其相似，其特征是有一个宽阔的、有点圆柱形的 antrum，紧接着在导管囊的前方有一个明显弯曲的区域，在左侧形成一个弱的憩室，以及一个呈小箭头形硬化体斑块形式的交配痕，通常有一个浅袋。雄性生殖器形态与新美苔蛾类群（蛀果蛾族）有许多相似之处，特别是Afroploce Aarvik。Gibbalaria的单系性得到以下特征的支持：两性前翅上凸起、卷曲、乳白色的鳞片，这在亚科中其他一些分散的属中也有出现（例如，Grapholitini族中的Thylacandra Diakonoff；Olethreutini族中的Astronauta Diakonoff和Cosmopoda Diakonoff）；雌性生殖器导管囊中的角状憩室；从瓣片下缘紧挨着兜基部长出的近矩形凸缘；以及在基部腔上方的刚毛状突出脊，这似乎与新美苔蛾类群许多属中瓣片的亚基部突起同源。此外，Diakonoffiana graziani Razowski，2015（模式产地：喀麦隆共和国）被转移到Cosmopoda Diakonoff，成为C. graziani（Razowski），新组合。
