Simpson Michael G, York Dana A
Department of Biology, San Diego State University, San Diego, California 92182, USA San Diego State University San Diego United States of America.
Botany Research Associate, California Academy of Sciences (CAS), 55 Music Concourse Drive, Golden Gate Park, San Francisco, California 94118, USA California Academy of Sciences San Francisco United States of America.
PhytoKeys. 2024 Oct 15;247:155-172. doi: 10.3897/phytokeys.247.132060. eCollection 2024.
D.A.York & M.G.Simpson (Boraginaceae) is described as new. This species is currently known to occur in serpentine barrens in the Shasta-Trinity National Forest of Siskiyou County, California, with one outlier population in possible serpentine of Lake County, California. The new species is most similar to and to , these three likely each others' closest relatives. All three have a relatively large corolla limb width and similar smooth, lance-ovate to ovate, marginally rounded, acuminate and abaxially transversely flattened nutlets. differs from in having a short, as opposed to a tall, stem height; bifurcate as opposed to trifurcate primary axis cymules; and typically 2-3 nutlets per fruit, as opposed to usually one nutlet per fruit. differs from also in having a short, versus tall, stem height; appressed-strigose and spreading-hispid stem vestiture, as opposed to strigose only or strigose and hirsute; calyx trichomes with two distinct vestiture types, these marginally appressed hirsute and medially hispid, as opposed to calyx trichomes of one type, dense, appressed to ascending, whitish sericeous; and 2-3 nutlets per fruit, as opposed to one nutlet per fruit. is relatively rare and joins seven other species that are found on serpentine, either obligately or facultatively. Current molecular phylogenetic studies support the mostly convergent evolution of serpentine adaptation in , but additional studies are needed.