Shavrin Alexey V
Daugavpils University; Institute of Life Sciences and Technologies; Coleopterological Research Centre; Vienibas 13; Daugavpils; LV-5401; Latvia.
Zootaxa. 2024 Oct 30;5531(1):1-62. doi: 10.11646/zootaxa.5531.1.1.
Five genera of the "Hygrogeus" complex of the omaliine tribe Anthophagini Thomson, 1859 are (re-)described and illustrated: Hygrogeus Mulsant & Rey, 1880, Trichodromeus Luze, 1903, Asiageodromicus gen. n., Himalodromeus gen. n. and Microhygrodromicus gen. n. The genus Hygrogeus contains three species: H. aemulus (Rosenhauer, 1847), known from the Middle European Alps, and two species with uncertain generic status from Kashmir: H. minor Coiffait, 1983 and H. rugosicollis Coiffait, 1983. The genus Trichodromeus contains 21 species distributed in the eastern Palaearctic Region. Four species of Trichodromeus are (re-)described and illustrated: T. kumaonensis (Champion, 1925) (India), T. ketmeniensis (Bordoni, 1985) (Kazakhstan, Kyrgyzstan, Turkmenistan), T. penicillatus (Reitter, 1900) (Kazakhstan, Uzbekistan, Kyrgyzstan, Tajikistan), and T. telnovi sp. n. (Nepal). A new synonymy is established: T. ketmeniensis = T. ovchinnikovi Shavrin & Klimenko, 2008 syn. n. Eight new combinations are proposed: T. alticola (Coiffait, 1983) comb. n. (from Hygrogeus), T. chinensis (Cheng & Peng, 2019) comb. n. (from Hygrogeus), T. gulmargensis (Coiffait, 1983) comb. n. (from Hygrogeus), T. languidus (Coiffait, 1983) comb. n. (from Hygrogeus), T. longipalpis Coiffait, 1983 comb. n. (from Geodromicus Redtenbacher, 1857), T. meurguesae Coiffait, 1979 comb. n. (from Hygrogeus), T. montanus (Coiffait, 1983) comb. n. (from Hygrogeus), and T. sulcicollis (Coiffait, 1983) comb. n. (from Hygrogeus). The habitus of T. pusillus (Coiffait, 1983), and habitus and aedeagi of T. armatus (Cameron, 1941) and T. jaegeri Shavrin, 2021 are illustrated. The genus Asiageodromicus gen. n., distributed mainly in the Himalayan Region and Tibet, contains nine species. Eight species of them were previously attributed to the crassipalpis group (Shavrin 2019) of Geodromicus: A. amplissimus (Shavrin, 2019) comb. n., A. crassipalpis (Champion, 1920) comb. n., A. dolpoensis (Coiffait, 1982) comb. n., A. namucoicus (Cheng & Peng, 2020) comb. n., A. palpalis (Coiffait, 1977) comb. n., A. spadiceus (Shavrin, 2019) comb. n., A. subquadratus (Cheng & Peng, 2020) comb. n., and A. trapezipennis (Coiffait, 1979) comb. n. Two species are (re-)described and illustrated: A. dolpoensis and A. weiperti sp. n. (eastern Nepal). The Himalayan genus Himalodromeus gen. n. contains three species. All these species are (re-)described and illustrated: H. cyanipennis (Cameron, 1924) comb. n. (from Hygrogeus), known from India and Nepal, H. grandis sp. n. (Pakistan) and H. reductus sp. n. (Nepal). A new synonymy is proposed: H. cyanipennis = Hygrogeus coeruleipennis Coiffait, 1983 syn. n. Lectotypes for Hygrogeus cyanipennis Cameron, 1924 and Hygrogeus cyanipennis atrocaeruleus Champion, 1925 are designated. The genus Microhygrodromicus gen. n. contains two new species from Yunnan (China): M. exiguus sp. n. and M. politus sp. n. Keys for genera of the "Hygrogeus" complex, all species of Asiageodromicus gen. n., Himalodromeus gen. n. and Microhygrodromicus gen. n. are provided. Distributional maps of T. jaegeri Shavrin, 2021, T. ketmeniensis, T. penicillatus, T. telnovi sp. n., and all species of the new genera are given. Five species are recorded for the first time: A. crassipalpis, H. cyanipennis and T. jaegeri from Nepal, T. penicillatus from Uzbekistan and Kyrgyzstan, and T. pusillus from Pakistan.
对1859年汤姆森氏Anthophagini族奥马利尼科“湿栖甲”复合体的五个属进行了（重新）描述和图示：湿栖甲属Mulsant & Rey，1880；毛角甲属Trichodromeus Luze，1903；亚洲土栖甲属Asiageodromicus gen. n.；喜马拉雅土栖甲属Himalodromeus gen. n.；微湿栖甲属Microhygrodromicus gen. n.。湿栖甲属包含三个物种：中欧阿尔卑斯山已知的蓝湿栖甲H. aemulus（罗森豪尔，1847），以及克什米尔地区两个属地位不确定的物种：小湿栖甲H. minor科法伊特，1983和皱胸湿栖甲H. rugosicollis科法伊特，1983。毛角甲属包含21个物种，分布于古北区东部。对毛角甲属的四个物种进行了（重新）描述和图示：库蒙毛角甲T. kumaonensis（钱皮恩，1925）（印度）、克捷缅毛角甲T. ketmeniensis（博尔托尼，1985）（哈萨克斯坦、吉尔吉斯斯坦、土库曼斯坦）、帚毛角甲T. penicillatus（赖特尔，1900）（哈萨克斯坦、乌兹别克斯坦、吉尔吉斯斯坦、塔吉克斯坦）和特尔诺夫毛角甲T. telnovi sp. n.（尼泊尔）。建立了一个新异名：克捷缅毛角甲 = 奥夫钦尼科夫毛角甲T. ovchinnikovi沙夫林 & 克利缅科，2008 syn. n.。提出了八个新组合：高山毛角甲T. alticola（科法伊特，1983）comb. n.（来自湿栖甲属）、中华毛角甲T. chinensis（程 & 彭，2019）comb. n.（来自湿栖甲属）、古尔马尔格毛角甲T. gulmargensis（科法伊特，1983）comb. n.（来自湿栖甲属）、柔弱毛角甲T. languidus（科法伊特，1983）comb. n.（来自湿栖甲属）、长须毛角甲T. longipalpis科法伊特，1983 comb. n.（来自土栖甲属Redtenbacher，1857）、默尔盖毛角甲T. meurguesae科法伊特，1979 comb. n.（来自湿栖甲属）、山地毛角甲T. montanus（科法伊特，1983）comb. n.（来自湿栖甲属）和沟胸毛角甲T. sulcicollis（科法伊特，1983）comb. n.（来自湿栖甲属）。图示了短小毛角甲T. pusillus（科法伊特，1983）的外形，以及武装毛角甲T. armatus（卡梅伦，1941）和耶格毛角甲T. jaegeri沙夫林，2021的外形和阳茎。亚洲土栖甲属Asiageodromicus gen. n.主要分布于喜马拉雅地区和西藏，包含九个物种。其中八个物种先前被归入土栖甲属的粗须组（沙夫林，2019）：粗壮亚洲土栖甲A. amplissimus（沙夫林，2019）comb. n.、粗须亚洲土栖甲A. crassipalpis（钱皮恩，1920）comb. n.、多尔波亚洲土栖甲A. dolpoensis（科法伊特，1982）comb. n.、纳木错亚洲土栖甲A. namucoicus（程 & 彭，2020）comb. n.、须状亚洲土栖甲A. palpalis（科法伊特，1977）comb. n.、栗色亚洲土栖甲A. spadiceus（沙夫林，2019）comb. n.、近方形亚洲土栖甲A. subquadratus（程 & 彭，2020）comb. n.和梯形翅亚洲土栖甲A. trapezipennis（科法伊特，1979）comb. n.。对两个物种进行了（重新）描述和图示：多尔波亚洲土栖甲和魏珀特亚洲土栖甲A. weiperti sp. n.（尼泊尔东部）。喜马拉雅属喜马拉雅土栖甲属Himalodromeus gen. n.包含三个物种。对所有这些物种进行了（重新）描述和图示：蓝翅喜马拉雅土栖甲H. cyanipennis（卡梅伦，1924）comb. n.（来自湿栖甲属），已知分布于印度和尼泊尔，大喜马拉雅土栖甲H. grandis sp. n.（巴基斯坦）和缩小喜马拉雅土栖甲H. reductus sp. n.（尼泊尔）。提出了一个新异名：蓝翅喜马拉雅土栖甲 = 蓝翅湿栖甲Hygrogeus coeruleipennis科法伊特，1983 syn. n.。指定了1924年卡梅伦氏蓝翅湿栖甲和1925年钱皮恩氏蓝翅暗蓝湿栖甲的选模。微湿栖甲属Microhygrodromicus gen. n.包含来自中国云南的两个新物种：微小微湿栖甲M. exiguus sp. n.和优美微湿栖甲M. politus sp. n.。提供了“湿栖甲”复合体各属、亚洲土栖甲属Asiageodromicus gen. n.、喜马拉雅土栖甲属Himalodromeus gen. n.和微湿栖甲属Microhygrodromicus gen. n.所有物种的检索表。给出了2021年沙夫林氏耶格毛角甲、克捷缅毛角甲、帚毛角甲、特尔诺夫毛角甲以及新属所有物种的分布图。有五个物种首次被记录：尼泊尔的粗须亚洲土栖甲、蓝翅喜马拉雅土栖甲和耶格毛角甲，乌兹别克斯坦和吉尔吉斯斯坦的帚毛角甲，以及巴基斯坦的短小毛角甲。
