Berdyev A
Parazitologiia. 1985 Mar-Apr;19(2):134-8.
The burrow-nest life scheme of ixodid ticks, which was formed at the beginning of the Chalk, gave rise to the subsequent types of parasitism: pasture (end of the Chalk), pasture-burrow (end of Paleogene) and pasture-stall one (late Neogene). Changes in the climate in different geological epochs and regions together with the hosts' evolution, natural selection in the aggregate of variability and heredity of ixodids in their populations were the cause of the evolution.
硬蜱的洞穴巢穴生活模式形成于白垩纪初期,随后产生了后续的寄生类型:牧场型(白垩纪末期)、牧场洞穴型(古近纪末期)和牧场畜栏型(新近纪晚期)。不同地质时期和地区的气候变化,以及宿主的进化、硬蜱种群在变异性和遗传性总和中的自然选择,是进化的原因。