Relationship between trackmakers of the Laetoli footprints from gait synchronization.

The parallel trails of footprints at Laetoli site G are important fossils for studying the characteristics of . However, the relationship between the trackmakers - i.e. whether it was that of an adult male-female pair or of parent-offspring - remains unclear. The footprints show that the two individuals walked side by side with a narrow and constant distance between them and synchronized their leg movements and step lengths (gait synchronization), although they had a large height difference. In this study, live camera videos were collected to obtain data on gait synchronization in , the closest extant species to . The data showed that when two humans with a large height difference walked alongside each other, with (at least) one of the pair having their arm around the other's shoulder or back, adult male-female pairs (couples) frequently synchronized their gait, but parent-offspring pairs did not, whereas both couples and parent-offspring seldom synchronized when they walked side by side without connection or with handholding. Two individuals only maintained a narrow and constant distance like that between the Laetoli footprints when they walked with an arm-around connection. Therefore, assuming that had the same gait synchronization tendency as , the trackmakers were more likely to be an adult male-female pair than a parent-offspring one.