Herreid C F, Lee L W, Shah G M
Respir Physiol. 1979 May;37(1):109-20. doi: 10.1016/0034-5687(79)90096-3.
Land Crabs, Cardisoma guanhumi, were fitted with respiratory masks and E.C.G. electrodes and run for 10 or 20 min on a treadmill at speeds of 150 and 300 cm/min. Aerobic metabolism increased linearly with the speed of locomotion. The recovery period was characterized by a large oxygen debt. The primary respiratory adjustment to exercise was an increased ventilation volume; only a minor increase in oxygen extraction occurred. The respiratory exchange ratio increased during exercise and during recovery, presumably correlated with a metabolic acidosis. These results are similar to data collected for exercising vertebrates and the net cost of locomotion of crabs appears similar to quadrupeds. However, the heart rate in exercising crabs changed in an unexpected way: during moderate exercise no change was noted, but during heavy exercise a bradycardia developed. The reduction in rate resulted from an increase in interbeat interval and frequent pauses in the heart beat.
将陆地蟹（关氏硬壳蟹）装上呼吸面罩和心电图电极，在跑步机上以150和300厘米/分钟的速度奔跑10或20分钟。有氧代谢随运动速度呈线性增加。恢复期的特点是存在大量氧债。对运动的主要呼吸调节是通气量增加；氧摄取仅略有增加。呼吸交换率在运动期间和恢复期间增加，推测与代谢性酸中毒有关。这些结果与对运动脊椎动物收集的数据相似，蟹类运动的净成本似乎与四足动物相似。然而，运动蟹的心率变化出人意料：在适度运动期间未观察到变化，但在剧烈运动期间出现心动过缓。心率降低是由于心跳间期延长和心跳频繁停顿所致。
