Barel C D, van der Meulen J W, Berkhoudt H
Anat Anz. 1977;142(1-2):21-31.
Through chains of ligaments and bones, which can be reduced to four-bar-systems (fbs), contraction of the sternohyoid and of the opercular levator muscle result in mandibular depression. Comparatively the opercular fbs transmits movement more efficiently than the hyoid fbs which, consequently, transmits force more efficiently.
通过韧带和骨骼链（可简化为四杆系统，fbs），胸骨舌骨肌和鳃盖提肌的收缩会导致下颌下降。相比之下，鳃盖四杆系统比舌骨四杆系统更有效地传递运动，因此，舌骨四杆系统更有效地传递力量。
