Roshdy M A, Hoogstraal H, Clifford C M, Keirans J E, Corwin M D
J Parasitol. 1984 Jun;70(3):407-11.
Adults of 4 of the 6 species constituting the subgenus Carios and of 3 of the 4 species constituting the subgenus Chiropterargas were studied by scanning electron microscopy. All species parasitize Old World cave-dwelling insectivorous bats (Microchiroptera). The anterior pit setae number 10 in Carios and 10 or 11 in Chiropterargas. In most Carios, the setiform seta is replaced by a second serrate seta. In 2 of the 3 studied Chiropterargas species, 1 of the 2 grooved setae is exceptionally long. Porose setae number 3 in Carios and 3 or 4 in Chiropterargas. The Haller's organ roof in both subgenera is solid, lacking perforations; the aperture is narrowly transverse in Carios, irregularly wide or wide and transverse in Chiropterargas; uniquely, 1 or 2 sensilla protrude from the aperture of Chiropterargas species. The protruding sensilla and long grooved seta of Chiropterargas suggest a probably distinctive sensory-behavior pattern common to these ticks. Other morphological characters are discussed and compared to show relationships between these 2 subgenera and the subgenera Argas and Persicargas and distinctive characters present only in adult and/or larval Carios and Chiropterargas.
通过扫描电子显微镜对构成卡里奥斯亚属的6个物种中的4个物种的成虫以及构成翼手argas亚属的4个物种中的3个物种的成虫进行了研究。所有物种均寄生于东半球洞穴栖息的食虫蝙蝠（小蝙蝠亚目）。卡里奥斯的前坑刚毛数量为10根，翼手argas的前坑刚毛数量为10根或11根。在大多数卡里奥斯物种中，刚毛状刚毛被第二根锯齿状刚毛取代。在所研究的翼手argas物种的3个物种中的2个物种中，2根带槽刚毛中的1根异常长。卡里奥斯的多孔刚毛数量为3根，翼手argas的多孔刚毛数量为3根或4根。两个亚属的哈勒氏器顶部都是实心的，没有穿孔；卡里奥斯的开口呈狭窄的横向，翼手argas的开口不规则地宽或宽且横向；独特的是，1根或2根感觉器从翼手argas物种的开口处突出。翼手argas的突出感觉器和长带槽刚毛表明这些蜱可能具有独特的感觉行为模式。讨论并比较了其他形态特征，以显示这两个亚属与argas亚属和波斯argas亚属之间的关系，以及仅在成年和/或幼虫卡里奥斯和翼手argas中出现的独特特征。
