Jouin C
Tissue Cell. 1978;10(2):289-301. doi: 10.1016/0040-8166(78)90024-1.
The epidermal and stomodeal cuticles of Protodrilus are described then compared. The thin epidermal cuticle, the thickness of which is about the same over all the body, is characterized both by the absence of fibrils in its deepest part and by the extension of epidermal microvilli above the cuticle. The stomodeal cuticle, the thickness of which is as variable as that of the epithelium, presents two layers of fibrils comparable to the collagen fibrils described in the cuticle of other Annelida, as well as a relatively diversified supramicrovillous coating. The anterior cuticular thickening or grating plate, is characterized by the length of the epithelial microvilli, the thickness of the cuticular matrix and the superficial cuticular zone with supramicrovillous denticles supported by an axis of fibrous bundles. In the stomodeal cuticle, the fibrillar material seems to give to the cuticle best resistance to deformation during the pharyngeal bulb contraction, while an especially elaborated supramicrovillous coating is found in regions most exposed to friction. These features contrast with the relative simplicity of the epidermal cuticle.
对原蚓的表皮角质层和口道角质层进行了描述，然后进行了比较。薄的表皮角质层在整个身体上厚度大致相同，其特征在于最深层没有纤维，且表皮微绒毛在角质层上方延伸。口道角质层的厚度与上皮层一样多变，呈现出两层与其他环节动物角质层中描述的胶原纤维相当的纤维，以及相对多样化的超微绒毛涂层。前部角质层增厚或格栅板的特征在于上皮微绒毛的长度、角质层基质的厚度以及由纤维束轴支撑的带有超微绒毛小齿的表面角质层区域。在口道角质层中，纤维状物质似乎使角质层在咽球收缩期间对变形具有最佳抵抗力，而在最易受摩擦的区域发现了特别精细的超微绒毛涂层。这些特征与表皮角质层的相对简单形成对比。
