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复杂不可逆性与进化

Complex-irreversibility and evolution.

作者信息

Walker I

出版信息

Experientia. 1983 Aug 15;39(8):806-13. doi: 10.1007/BF01990396.

Abstract

Both, irreversibility and evolution, imply order in time. It is argued that the only possible concept of time is a 'system-specific time', and that order in time is convertible into order in space and vice-versa. While life-less, complex systems are irreversible because of their complexity and, hence, not repeatable, living systems are reproduced by irreversible copy-reproduction and by coding. This mode of reproduction results of necessity in an arrow of time of growth and increasing complexity with death as its antagonist, and in obligatory spatial asymmetry. This arrow of increasing organic complexity is simultaneous with, and independent of, the arrow of increasing entropy. - A generalized, organic hierarchy is proposed as the model to study higher evolution. This hierarchy reproduces itself by differential rates of reproduction of its subunits within and between the various hierarchical levels of organization. Phylogenetic change is brought about by a change in this hierarchy's specific phase pattern of growth. Continuous and discrete organization is defined, and it is shown that specific relations between continuous and discrete levels within the hierarchy result in accumulation of neutral alleles. This accumulation is due to complex-irreversibility and causes genetic stabilisation, i.e. heritability, of the species-specific morphology of organisms.

摘要

不可逆性和进化都意味着时间上的有序性。有人认为,唯一可能的时间概念是“系统特定时间”,并且时间上的有序性可以转化为空间上的有序性,反之亦然。无生命的复杂系统由于其复杂性而具有不可逆性,因此不可重复,而生命系统则通过不可逆的复制繁殖和编码进行繁殖。这种繁殖方式必然导致以死亡为对立面的生长和复杂性增加的时间箭头,以及必然的空间不对称性。这种有机复杂性增加的箭头与熵增加的箭头同时存在且相互独立。——提出了一个广义的有机层次结构作为研究高级进化的模型。这个层次结构通过其亚单位在组织的各个层次内部和之间不同的繁殖速率来自我复制。系统发育变化是由这个层次结构特定的生长阶段模式的变化引起的。定义了连续组织和离散组织,并表明层次结构中连续和离散层次之间的特定关系导致中性等位基因的积累。这种积累是由于复杂不可逆性,导致生物体物种特异性形态的遗传稳定,即遗传性。

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