Michel K J, Petersen M
Cell Tissue Res. 1982;222(3):667-76. doi: 10.1007/BF00213864.
The tympanal organ of the migratory locust acquires its definitive form during larval development. All the receptor cells (90-100) are present in the 1st instar, whereas the differentiation of the tympanum and the cuticular structures it bears proceeds in steps from one instar to the next. The elevated process is the earliest such structure to appear (2nd instar); it is followed by the pyriform vesicle (3rd instar) and folded body (4th instar). The styliform body first appears in the imago. Although the typical arrangement of the receptor cells is already discernible in the 1st instar, some of the attachment sites change during development, the final configuration appearing only in the imago.
飞蝗的鼓膜听器在幼虫发育过程中形成其最终形态。所有受体细胞（90 - 100个）在一龄幼虫时就已存在，而鼓膜及其所承载的表皮结构的分化是从一个龄期到下一个龄期逐步进行的。隆起过程是最早出现的此类结构（二龄幼虫）；随后是梨形囊泡（三龄幼虫）和折叠体（四龄幼虫）。柱状体首先出现在成虫中。尽管受体细胞的典型排列在一龄幼虫时就已可辨，但一些附着位点在发育过程中会发生变化，最终构型仅在成虫中出现。
