Klag J
J Embryol Exp Morphol. 1982 Dec;72:183-95.
In the early embryo of Tetrodontophora bielanensis (up to the stage of 500 blastomeres) nuage granules occur in two different locations: (1) in areas where the invaginating cleavage furrows have pushed fragments of the oosome into the yolk mass, and (2) in the oosome proper. In the first areas the granules are few in number and certain cells that have enclosed them in their cytoplasm eventually degenerate. The remaining cells arising in these areas are devoid of any nuage granules and differentiate into yolk cells. A different situation is observed in the other areas, where certain cells resulting from tangential divisions of the superficial blastomeres contain many nuage granules and represent primordial germ cells (PGCs). The incipient PGCs differ from the other cells of the embryo in possessing nuage granules associated with mitochondria and in lacking any annulate lamellae.
在双齿多刺蚁的早期胚胎中（直至500个卵裂球阶段），生殖质颗粒出现在两个不同的位置：（1）在内陷的卵裂沟将卵质体碎片挤入卵黄块的区域，以及（2）在卵质体本身。在第一个区域，颗粒数量很少，某些将它们包裹在细胞质中的细胞最终会退化。在这些区域产生的其余细胞没有任何生殖质颗粒，并分化为卵黄细胞。在其他区域观察到不同的情况，在这些区域，由表面卵裂球的切向分裂产生的某些细胞含有许多生殖质颗粒，并代表原始生殖细胞（PGC）。初始的PGC与胚胎的其他细胞不同，它们具有与线粒体相关的生殖质颗粒，并且没有任何环形片层。
